%PDF-1.6
%
1 0 obj
<>
endobj
2 0 obj
<>stream
C. Melissa Miner, Jennifer L. Burnaford, Richard F. Ambrose, Liam Antrim, Heath Bohlmann, Carol A. Blanchette, John M. Engle, Steven C. Fradkin, Rani Gaddam, Christopher D. G. Harley, Benjamin G. Miner, Steven N. Murray, Jayson R. Smith, Stephen G. Whitaker, Peter T. Raimondi
Large-scale impacts of sea star wasting disease (SSWD) on intertidal sea stars and implications for recovery
www.plosone.org
www.plosone.org
endstream
endobj
3 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 12 0 R/Annots 13 0 R/Contents 14 0 R/TrimBox[0 0 612 792]>>
endobj
13 0 obj
[15 0 R 16 0 R 17 0 R 18 0 R 19 0 R 20 0 R 21 0 R 22 0 R 23 0 R 24 0 R 25 0 R 26 0 R 27 0 R 28 0 R 29 0 R 30 0 R 31 0 R 32 0 R 33 0 R 34 0 R 35 0 R 36 0 R 37 0 R 38 0 R 39 0 R 40 0 R 41 0 R 42 0 R 43 0 R 44 0 R 45 0 R 46 0 R 47 0 R 48 0 R 49 0 R]
endobj
15 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref001)>>
endobj
16 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref003)>>
endobj
17 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref002)>>
endobj
18 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref004)>>
endobj
19 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref005)>>
endobj
20 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref006)>>
endobj
21 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref007)>>
endobj
22 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref008)>>
endobj
23 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref009)>>
endobj
24 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref010)>>
endobj
25 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref011)>>
endobj
26 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref008)>>
endobj
27 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref012)>>
endobj
28 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref015)>>
endobj
29 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref016)>>
endobj
30 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref007)>>
endobj
31 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref017)>>
endobj
32 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref018)>>
endobj
33 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref007)>>
endobj
34 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref007)>>
endobj
35 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref019)>>
endobj
36 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref017)>>
endobj
37 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref019)>>
endobj
38 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref012)>>
endobj
39 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref012)>>
endobj
40 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref014)>>
endobj
41 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref020)>>
endobj
42 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref012)>>
endobj
43 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref014)>>
endobj
44 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref020)>>
endobj
45 0 obj
<>/Border[0 0 0]/A 50 0 R>>
endobj
50 0 obj
<>
endobj
46 0 obj
<>/Border[0 0 0]/A 51 0 R>>
endobj
51 0 obj
<>
endobj
47 0 obj
<>/Border[0 0 0]/A 52 0 R>>
endobj
52 0 obj
<>
endobj
48 0 obj
<>/Border[0 0 0]/A 53 0 R>>
endobj
53 0 obj
<>
endobj
49 0 obj
<>/Border[0 0 0]/A 54 0 R>>
endobj
54 0 obj
<>
endobj
14 0 obj
[55 0 R 56 0 R 57 0 R 58 0 R 59 0 R 60 0 R 61 0 R 62 0 R 63 0 R 64 0 R 65 0 R 66 0 R 67 0 R 68 0 R 69 0 R 70 0 R 71 0 R 72 0 R 73 0 R 74 0 R 75 0 R]
endobj
55 0 obj
<>stream
q
0.45 0.36 0.35 0.01 k
0 w
575.9997 624.2298 m
199.9998 624.2298 l
199.9998 624.7298 l
575.9997 624.7298 l
f*
0.83 0.64 0.02 0 k
517.1528 549.9213 m
521.915 549.9213 l
h
f*
527.074 549.9213 m
531.8362 549.9213 l
h
f*
504.5102 510.9165 m
509.2157 510.9165 l
h
f*
511.4835 510.9165 m
516.2457 510.9165 l
h
f*
370.4882 497.9339 m
375.2504 497.9339 l
h
f*
466.2992 497.9339 m
471.0614 497.9339 l
h
f*
558.3685 497.9339 m
563.1307 497.9339 l
h
f*
494.7024 458.9291 m
499.4646 458.9291 l
h
f*
382.9606 432.9071 m
387.7228 432.9071 l
h
f*
203.4142 419.9244 m
212.9386 419.9244 l
h
f*
522.3685 419.9244 m
531.8929 419.9244 l
h
f*
459.4961 380.9197 m
464.2583 380.9197 l
h
f*
466.526 380.9197 m
475.9937 380.9197 l
h
f*
481.2094 380.9197 m
490.6772 380.9197 l
h
f*
280.1197 341.915 m
289.6441 341.915 l
h
f*
362.6079 315.9496 m
367.3701 315.9496 l
h
f*
369.6378 315.9496 m
379.1622 315.9496 l
h
f*
384.3213 315.9496 m
393.8457 315.9496 l
h
f*
319.2378 302.9102 m
324 302.9102 l
h
f*
334.5449 289.9276 m
339.3071 289.9276 l
h
f*
341.5181 289.9276 m
351.0425 289.9276 l
h
f*
364.5921 276.9449 m
374.1165 276.9449 l
h
f*
376.3843 276.9449 m
385.852 276.9449 l
h
f*
382.1669 224.9008 m
391.6913 224.9008 l
h
f*
346.7906 198.9354 m
356.315 198.9354 l
h
f*
361.474 198.9354 m
370.9984 198.9354 l
h
f*
373.2661 198.9354 m
382.7906 198.9354 l
h
f*
459.2126 146.948 m
468.737 146.948 l
h
f*
473.8961 146.948 m
483.4205 146.948 l
h
f*
487.8992 146.948 m
497.3669 146.948 l
h
f*
0 g
1 j
1 J
9.5 0 0 9.5 200.0125 707.7543 cm
BT
/F0 1 Tf
1 TL
-0.0042 Tc
0 0 Td
(than)Tj
2.1423 0 Td
(during)Tj
2.96 0 Td
(pre-SSWD)Tj
4.9651 0 Td
(periods.)Tj
3.7298 0 Td
(In)Tj
1.0503 0 Td
(hindsight,)Tj
4.4459 0 Td
(our)Tj
1.6471 0 Td
(data)Tj
2.1423 0 Td
(suggest)Tj
3.6821 0 Td
(that)Tj
1.8738 0 Td
(the)Tj
1.5934 0 Td
(SSWD)Tj
3.2106 0 Td
(event)Tj
2.6377 0 Td
(defied)Tj
-36.0804 -1.474 Td
(prediction)Tj
4.4936 0 Td
(based)Tj
2.9122 0 Td
(on)Tj
1.3189 0 Td
(two)Tj
1.7604 0 Td
(factors)Tj
3.1987 0 Td
(found)Tj
2.6914 0 Td
(to)Tj
1.0443 0 Td
(be)Tj
1.3189 0 Td
(important)Tj
4.3444 0 Td
(in)Tj
0.9847 0 Td
(other)Tj
2.4706 0 Td
(marine)Tj
3.2404 0 Td
(disease)Tj
3.6224 0 Td
(events,)Tj
3.4135 0 Td
(sea)Tj
-36.8144 -1.474 Td
(water)Tj
2.6377 0 Td
(temperature)Tj
5.5558 0 Td
(and)Tj
1.8679 0 Td
(population)Tj
4.7682 0 Td
(density,)Tj
3.6283 0 Td
(and)Tj
1.8679 0 Td
(illustrate)Tj
3.8909 0 Td
(the)Tj
1.5934 0 Td
(importance)Tj
5.1142 0 Td
(of)Tj
1.0444 0 Td
(surveillance)Tj
5.4305 0 Td
(of)Tj
-37.3992 -1.474 Td
(natural)Tj
3.2344 0 Td
(populations)Tj
5.2635 0 Td
(as)Tj
1.2651 0 Td
(one)Tj
1.8679 0 Td
(element)Tj
3.7358 0 Td
(of)Tj
1.0503 0 Td
(an)Tj
1.3188 0 Td
(integrated)Tj
4.6071 0 Td
(approach)Tj
4.3384 0 Td
(to)Tj
1.0503 0 Td
(marine)Tj
3.2405 0 Td
(disease)Tj
3.6223 0 Td
(ecology.)Tj
-34.5944 -1.474 Td
(Low)Tj
2.029 0 Td
(levels)Tj
2.7451 0 Td
(of)Tj
1.0443 0 Td
(SSWD-symptoma)Tj
7.8952 0 Td
(tic)Tj
1.2115 0 Td
(sea)Tj
1.8082 0 Td
(stars)Tj
2.3691 0 Td
(are)Tj
1.6471 0 Td
(still)Tj
1.6411 0 Td
(present)Tj
3.5209 0 Td
(throughout)Tj
4.9412 0 Td
(the)Tj
1.5994 0 Td
(impacted)Tj
4.231 0 Td
(range,)Tj
-36.6831 -1.474 Td
(thus)Tj
2.0886 0 Td
(the)Tj
1.5994 0 Td
(outlook)Tj
3.4015 0 Td
(for)Tj
1.3726 0 Td
(population)Tj
4.7681 0 Td
(recovery)Tj
4.0163 0 Td
(is)Tj
0.9309 0 Td
(uncertain.)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 568.0062 cm
BT
/F2 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Introduction)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 550.9984 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(Infectious)Tj
4.2009 0 Td
(diseases)Tj
3.3788 0 Td
(are)Tj
1.4514 0 Td
(prevalent)Tj
3.9344 0 Td
(in)Tj
1.0375 0 Td
(terrestrial)Tj
4.0875 0 Td
(and)Tj
1.7292 0 Td
(marine)Tj
3.084 0 Td
(systems)Tj
3.2995 0 Td
(\(reviewed)Tj
4.1386 0 Td
(in)Tj
1.0318 0 Td
([)Tj
0.83 0.64 0.02 0 k
(1)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(3)Tj
0 g
(]\),)Tj
2.9367 0 Td
(and)Tj
1.7291 0 Td
(can)Tj
-36.0394 -1.2982 Td
(result)Tj
2.4491 0 Td
(in)Tj
1.0318 0 Td
(mass)Tj
2.1996 0 Td
(mortality)Tj
3.9118 0 Td
(of)Tj
1.0148 0 Td
(a)Tj
0.6577 0 Td
(species)Tj
2.9933 0 Td
(or)Tj
1.0999 0 Td
(suite)Tj
2.0976 0 Td
(of)Tj
1.0148 0 Td
(species.)Tj
3.2201 0 Td
(Evidence)Tj
3.8608 0 Td
(suggests)Tj
3.4809 0 Td
(that)Tj
1.7802 0 Td
(disease)Tj
3.016 0 Td
(out-)Tj
-33.8284 -1.3039 Td
(breaks)Tj
2.8119 0 Td
(in)Tj
1.0375 0 Td
(marine)Tj
3.084 0 Td
(systems)Tj
3.2996 0 Td
(are)Tj
1.4456 0 Td
(becoming)Tj
4.1839 0 Td
(increasingly)Tj
5.0287 0 Td
(common,)Tj
4.0705 0 Td
(potentially)Tj
4.4617 0 Td
(as)Tj
1.0148 0 Td
(a)Tj
0.6519 0 Td
(consequence)Tj
5.3405 0 Td
(of)Tj
-36.4306 -1.2983 Td
(the)Tj
1.4683 0 Td
(shift)Tj
1.9615 0 Td
(in)Tj
1.0375 0 Td
(environmental)Tj
6.1115 0 Td
(conditions)Tj
4.473 0 Td
(associated)Tj
4.2576 0 Td
(with)Tj
1.9956 0 Td
(global)Tj
2.6249 0 Td
(climate)Tj
3.1351 0 Td
(change)Tj
3.0444 0 Td
([)Tj
0.83 0.64 0.02 0 k
(2)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(4)Tj
0 g
(].)Tj
2.3017 0 Td
(When)Tj
2.6815 0 Td
(a)Tj
0.6577 0 Td
(dis-)Tj
-35.7503 -1.2983 Td
(ease)Tj
1.8595 0 Td
(outbreak)Tj
3.787 0 Td
(affects)Tj
2.7326 0 Td
(a)Tj
0.6576 0 Td
(foundation)Tj
4.6715 0 Td
(species)Tj
2.999 0 Td
([)Tj
0.83 0.64 0.02 0 k
(5)Tj
0 g
(],)Tj
1.5988 0 Td
(ecosystem)Tj
4.2916 0 Td
(engineer)Tj
3.6907 0 Td
([)Tj
0.83 0.64 0.02 0 k
(6)Tj
0 g
(])Tj
1.3776 0 Td
(or)Tj
1.0999 0 Td
(keystone)Tj
3.7303 0 Td
(species)Tj
2.9991 0 Td
([)Tj
0.83 0.64 0.02 0 k
(7)Tj
0 g
(],)Tj
-35.4952 -1.3039 Td
(disease-induced)Tj
6.6103 0 Td
(changes)Tj
3.4072 0 Td
(in)Tj
1.0318 0 Td
(abundance,)Tj
4.8246 0 Td
(size)Tj
1.6837 0 Td
(structure,)Tj
4.0479 0 Td
(and)Tj
1.7291 0 Td
(individual)Tj
4.286 0 Td
(behavior)Tj
3.7133 0 Td
(and)Tj
1.7291 0 Td
(perfor-)Tj
-33.063 -1.2982 Td
(mance)Tj
2.8686 0 Td
(can)Tj
1.6271 0 Td
(ripple)Tj
2.5625 0 Td
(through)Tj
3.4412 0 Td
(the)Tj
1.474 0 Td
(entire)Tj
2.5512 0 Td
(ecosystem,)Tj
4.5184 0 Td
(causing)Tj
3.2428 0 Td
(substantial)Tj
4.5071 0 Td
(and)Tj
1.7291 0 Td
(long-term)Tj
4.2519 0 Td
(changes)Tj
3.4016 0 Td
(in)Tj
-36.1755 -1.2983 Td
(system)Tj
2.9366 0 Td
(structure)Tj
3.8211 0 Td
(and)Tj
1.7348 0 Td
(function.)Tj
3.8494 0 Td
(Examples)Tj
4.0592 0 Td
(of)Tj
1.0148 0 Td
(`marine)Tj
3.3108 0 Td
(disease)Tj
3.0161 0 Td
(emergencies')Tj
5.3858 0 Td
([)Tj
0.83 0.64 0.02 0 k
(8)Tj
0 g
(])Tj
1.3776 0 Td
(in)Tj
1.0318 0 Td
(which)Tj
2.6532 0 Td
(ecolog-)Tj
-34.1912 -1.2983 Td
(ically)Tj
2.2903 0 Td
(critical)Tj
2.9537 0 Td
(species)Tj
2.9934 0 Td
(have)Tj
2.0579 0 Td
(been)Tj
2.126 0 Td
(affected)Tj
3.3278 0 Td
(by)Tj
1.1792 0 Td
(disease)Tj
3.0161 0 Td
(outbreaks)Tj
4.1499 0 Td
(and)Tj
1.7291 0 Td
(subsequent)Tj
4.7055 0 Td
(mass)Tj
2.194 0 Td
(mortalities)Tj
-32.7229 -1.3039 Td
(include)Tj
3.1804 0 Td
(eelgrass)Tj
3.3108 0 Td
(wasting)Tj
3.2769 0 Td
(disease)Tj
3.016 0 Td
(\(reviewed)Tj
4.1386 0 Td
(in)Tj
1.0318 0 Td
([)Tj
0.83 0.64 0.02 0 k
(9)Tj
0 g
(]\),)Tj
1.9445 0 Td
(white-band)Tj
4.7849 0 Td
(disease)Tj
3.0217 0 Td
(in)Tj
1.0318 0 Td
(reef-building)Tj
5.4368 0 Td
(corals)Tj
-34.1742 -1.2983 Td
([)Tj
0.83 0.64 0.02 0 k
(10)Tj
0 g
(],)Tj
2.0749 0 Td
(an)Tj
1.2019 0 Td
(unidentified)Tj
5.1363 0 Td
(pathogen)Tj
3.9402 0 Td
(affecting)Tj
3.6566 0 Td
(the)Tj
1.4684 0 Td
(tropical)Tj
3.2881 0 Td
(urchin)Tj
/F10 1 Tf
2.8857 0 Td
[()-4()4()6()4()2()6()]TJ
3.8211 0 Td
[()16()22()7()4( )4( )4()16(\n)-9()12()]TJ
/F6 1 Tf
4.422 0 Td
([)Tj
0.83 0.64 0.02 0 k
(11)Tj
0 g
(])Tj
1.8538 0 Td
(and,)Tj
1.9503 0 Td
(as)Tj
-35.6993 -1.2982 Td
(reported)Tj
3.6566 0 Td
(here,)Tj
2.194 0 Td
(sea)Tj
1.44 0 Td
(star)Tj
1.6838 0 Td
(wasting)Tj
3.2768 0 Td
(disease)Tj
3.0161 0 Td
(\(SSWD\))Tj
3.5546 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4683 0 Td
(ochre)Tj
2.4718 0 Td
(sea)Tj
1.44 0 Td
(star)Tj
/F10 1 Tf
1.6837 0 Td
[()2()-4(\r)-3(\r)9()11()2(\n)]TJ
3.2315 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
(.)Tj
-28.9585 -1.3039 Td
(SSWD)Tj
2.8743 0 Td
(is)Tj
0.8504 0 Td
(an)Tj
1.2018 0 Td
(ongoing)Tj
3.5263 0 Td
(disease)Tj
3.0217 0 Td
(epidemic,)Tj
4.1102 0 Td
(which)Tj
2.6533 0 Td
(has)Tj
1.542 0 Td
(devastated)Tj
4.405 0 Td
(intertidal)Tj
3.9005 0 Td
(and)Tj
1.7291 0 Td
(nearshore)Tj
4.1896 0 Td
(sea)Tj
-35.2004 -1.2983 Td
(star)Tj
1.6837 0 Td
(populations)Tj
4.9493 0 Td
(along)Tj
2.4208 0 Td
(much)Tj
2.5228 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4683 0 Td
(west)Tj
1.9842 0 Td
(coast)Tj
2.2451 0 Td
(of)Tj
1.0148 0 Td
(North)Tj
2.6588 0 Td
(America)Tj
3.6454 0 Td
([)Tj
0.83 0.64 0.02 0 k
(8)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(12)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(15)Tj
0 g
(].)Tj
/F10 1 Tf
4.2463 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0308 0 Td
(is)Tj
0.8504 0 Td
(one)Tj
-35.7276 -1.2983 Td
(of)Tj
1.0147 0 Td
(the)Tj
1.4684 0 Td
(most)Tj
2.2053 0 Td
(recognized)Tj
4.5921 0 Td
(species)Tj
2.9934 0 Td
(from)Tj
2.2053 0 Td
(the)Tj
1.474 0 Td
(intertidal)Tj
3.9005 0 Td
(and)Tj
1.7291 0 Td
(shallow)Tj
3.2258 0 Td
(subtidal)Tj
3.3902 0 Td
(zones)Tj
2.4718 0 Td
(along)Tj
2.4208 0 Td
(this)Tj
1.6781 0 Td
(coast,)Tj
-34.7695 -1.3039 Td
(with)Tj
1.9955 0 Td
(a)Tj
0.652 0 Td
(broad)Tj
2.5682 0 Td
(geographic)Tj
4.6204 0 Td
(range)Tj
2.4661 0 Td
(from)Tj
2.2054 0 Td
(Prince)Tj
2.8119 0 Td
(William)Tj
3.4639 0 Td
(Sound)Tj
2.8063 0 Td
(Alaska,)Tj
3.1011 0 Td
(USA)Tj
2.1203 0 Td
(to)Tj
1.0261 0 Td
(Cedros)Tj
3.0728 0 Td
(Island,)Tj
2.9026 0 Td
(Baja)Tj
-35.8126 -1.2982 Td
(California,)Tj
4.473 0 Td
(Mexico)Tj
3.1975 0 Td
([)Tj
0.83 0.64 0.02 0 k
(16)Tj
0 g
(].)Tj
2.0749 0 Td
(It)Tj
0.8617 0 Td
(is)Tj
0.8447 0 Td
(a)Tj
0.652 0 Td
(generalist)Tj
4.0592 0 Td
(predator,)Tj
3.9004 0 Td
(with)Tj
1.9956 0 Td
(a)Tj
0.652 0 Td
(diet)Tj
1.7404 0 Td
(that)Tj
1.7745 0 Td
(includes)Tj
3.549 0 Td
(both)Tj
2.0579 0 Td
(sessile)Tj
2.6646 0 Td
(and)Tj
-34.4974 -1.2983 Td
(mobile)Tj
2.9876 0 Td
(invertebrate)Tj
5.04 0 Td
(prey)Tj
1.9899 0 Td
(items)Tj
2.3868 0 Td
(from)Tj
2.211 0 Td
(multiple)Tj
3.566 0 Td
(taxa)Tj
1.8481 0 Td
(\(including)Tj
4.3824 0 Td
(annelid)Tj
3.2144 0 Td
(worms,)Tj
3.1918 0 Td
(barnacles,)Tj
4.1896 0 Td
(bra-)Tj
-35.0076 -1.2983 Td
(chiopods,)Tj
4.0818 0 Td
(chitons,)Tj
3.3789 0 Td
(mussels,)Tj
3.5433 0 Td
(and)Tj
1.7348 0 Td
(whelks:)Tj
3.1804 0 Td
([)Tj
0.83 0.64 0.02 0 k
(7)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(17)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(18)Tj
0 g
(]\).)Tj
/F10 1 Tf
4.5864 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9978 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0309 0 Td
(was)Tj
1.6951 0 Td
(the)Tj
1.474 0 Td
(first)Tj
1.8028 0 Td
(species)Tj
2.9991 0 Td
(to)Tj
1.0261 0 Td
(be)Tj
-34.5314 -1.3039 Td
(termed)Tj
3.084 0 Td
(a)Tj
0.652 0 Td
(`keystone)Tj
3.9571 0 Td
(predator')Tj
3.8891 0 Td
([)Tj
0.83 0.64 0.02 0 k
(7)Tj
0 g
(])Tj
1.372 0 Td
(because)Tj
3.3222 0 Td
(of)Tj
1.0148 0 Td
(its)Tj
1.1452 0 Td
(strong)Tj
2.7722 0 Td
(effects)Tj
2.7269 0 Td
(on)Tj
1.2756 0 Td
(diversity)Tj
3.6454 0 Td
(of)Tj
1.0148 0 Td
(primary)Tj
3.4525 0 Td
(space)Tj
-33.3238 -1.2983 Td
(occupiers)Tj
4.0421 0 Td
(in)Tj
1.0318 0 Td
(wave-exposed)Tj
5.794 0 Td
(areas)Tj
2.245 0 Td
([)Tj
0.83 0.64 0.02 0 k
(7)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(19)Tj
0 g
(].)Tj
2.778 0 Td
(Even)Tj
2.2167 0 Td
(in)Tj
1.0318 0 Td
(areas)Tj
2.245 0 Td
(where)Tj
/F10 1 Tf
2.6532 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0309 0 Td
(does)Tj
2.0409 0 Td
(not)Tj
1.5704 0 Td
(play)Tj
1.8765 0 Td
(a)Tj
0.652 0 Td
(key-)Tj
-35.2004 -1.2982 Td
(stone)Tj
2.3584 0 Td
(role,)Tj
1.9899 0 Td
(it)Tj
0.788 0 Td
(is)Tj
0.8447 0 Td
(still)Tj
1.6441 0 Td
(a)Tj
0.652 0 Td
(dominant)Tj
4.1725 0 Td
(predator)Tj
3.668 0 Td
([)Tj
0.83 0.64 0.02 0 k
(17)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(19)Tj
0 g
(])Tj
3.0274 0 Td
(and)Tj
1.7291 0 Td
(thus)Tj
1.9389 0 Td
(is)Tj
0.8448 0 Td
(considered)Tj
4.6034 0 Td
(an)Tj
1.2019 0 Td
(ecologically)Tj
4.8642 0 Td
(impor-)Tj
-34.3273 -1.3039 Td
(tant)Tj
1.7971 0 Td
(species)Tj
2.999 0 Td
(throughout)Tj
4.7792 0 Td
(its)Tj
1.1452 0 Td
(range.)Tj
-9.5243 -1.2983 Td
(Determining)Tj
5.3914 0 Td
(the)Tj
1.4684 0 Td
(long-term)Tj
4.2519 0 Td
(effects)Tj
2.7269 0 Td
(of)Tj
1.0148 0 Td
(SSWD)Tj
2.8743 0 Td
(on)Tj
/F10 1 Tf
1.2756 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0309 0 Td
(populations)Tj
4.9492 0 Td
(and)Tj
1.7348 0 Td
(on)Tj
1.2756 0 Td
(rocky)Tj
-33.1821 -1.2983 Td
(intertidal)Tj
3.9004 0 Td
(communities)Tj
5.4822 0 Td
(poses)Tj
2.3924 0 Td
(a)Tj
0.652 0 Td
(particular)Tj
4.1102 0 Td
(challenge)Tj
3.9571 0 Td
(for)Tj
1.389 0 Td
(ecologists)Tj
4.0818 0 Td
(because)Tj
3.3166 0 Td
(the)Tj
1.4683 0 Td
(etiology)Tj
3.3902 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
-35.155 -1.3039 Td
(disease)Tj
3.016 0 Td
(is)Tj
0.8504 0 Td
(not)Tj
1.5704 0 Td
(yet)Tj
1.4003 0 Td
(fully)Tj
1.9785 0 Td
(resolved)Tj
3.5377 0 Td
(\(although)Tj
4.0932 0 Td
(see)Tj
1.4286 0 Td
([)Tj
0.83 0.64 0.02 0 k
(12)Tj
0 g
(])Tj
1.8482 0 Td
(for)Tj
1.389 0 Td
(identification)Tj
5.5842 0 Td
(of)Tj
1.0148 0 Td
(an)Tj
1.2019 0 Td
(associated)Tj
4.2576 0 Td
(denso-)Tj
-33.1708 -1.2983 Td
(virus\).)Tj
2.7665 0 Td
(Early)Tj
2.2961 0 Td
(signs)Tj
2.2223 0 Td
(of)Tj
1.0148 0 Td
(SSWD)Tj
2.8744 0 Td
(include)Tj
3.1804 0 Td
(a)Tj
0.6576 0 Td
(twisted)Tj
3.0784 0 Td
(or)Tj
1.0999 0 Td
(deflated)Tj
3.3902 0 Td
(appearance,)Tj
4.9719 0 Td
(followed)Tj
3.6454 0 Td
(by)Tj
1.1848 0 Td
(the)Tj
1.4684 0 Td
(develop-)Tj
-33.8511 -1.2982 Td
(ment)Tj
2.3073 0 Td
(of)Tj
1.0148 0 Td
(lesions)Tj
2.9367 0 Td
(\(authors')Tj
3.7984 0 Td
(pers.)Tj
2.1203 0 Td
(obs\);)Tj
2.16 0 Td
([)Tj
0.83 0.64 0.02 0 k
(12)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(14)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(20)Tj
0 g
(].)Tj
4.7225 0 Td
(Because)Tj
3.3959 0 Td
(these)Tj
2.2507 0 Td
(early)Tj
2.1486 0 Td
(symptoms)Tj
4.3597 0 Td
(are)Tj
1.4513 0 Td
(similar)Tj
2.982 0 Td
(to)Tj
-35.6482 -1.3039 Td
(those)Tj
2.3357 0 Td
(resulting)Tj
3.7304 0 Td
(from)Tj
2.2053 0 Td
(other)Tj
2.3471 0 Td
(sources)Tj
3.1974 0 Td
(of)Tj
1.0148 0 Td
(stress)Tj
2.3924 0 Td
(in)Tj
1.0375 0 Td
(sea)Tj
1.44 0 Td
(stars,)Tj
2.2677 0 Td
(such)Tj
2.0636 0 Td
(as)Tj
1.0148 0 Td
(desiccation)Tj
4.6998 0 Td
(or)Tj
1.0999 0 Td
(injury)Tj
2.6418 0 Td
(from)Tj
-33.4882 -1.2983 Td
(predators)Tj
4.0308 0 Td
(\(authors')Tj
3.8041 0 Td
(pers.)Tj
2.1146 0 Td
(obs.\),)Tj
2.3868 0 Td
(it)Tj
0.788 0 Td
(is)Tj
0.8447 0 Td
(impossible)Tj
4.4957 0 Td
(to)Tj
1.0262 0 Td
(know)Tj
2.4547 0 Td
(for)Tj
1.389 0 Td
(sure)Tj
1.9049 0 Td
(that)Tj
1.7745 0 Td
(SSWD)Tj
2.8743 0 Td
(is)Tj
0.8504 0 Td
(present)Tj
3.1634 0 Td
(in)Tj
1.0318 0 Td
(a)Tj
0.6576 0 Td
(pop-)Tj
-35.5915 -1.2983 Td
(ulation)Tj
3.05 0 Td
(until)Tj
2.1033 0 Td
(large)Tj
2.16 0 Td
(numbers)Tj
3.7814 0 Td
(of)Tj
1.0148 0 Td
(animals)Tj
3.3278 0 Td
(enter)Tj
2.2904 0 Td
(the)Tj
1.4683 0 Td
(later)Tj
1.99 0 Td
(stages)Tj
2.5625 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.474 0 Td
(disease)Tj
3.016 0 Td
(\(identified)Tj
4.3994 0 Td
(by)Tj
1.1792 0 Td
(large-)Tj
-34.8319 -1.2982 Td
(scale)Tj
2.1089 0 Td
(tissue)Tj
2.4548 0 Td
(degradation,)Tj
5.2497 0 Td
(body)Tj
2.2167 0 Td
(fragmentation,)Tj
6.1512 0 Td
(authors')Tj
3.4582 0 Td
(pers.)Tj
2.1203 0 Td
(obs;)Tj
1.8198 0 Td
([)Tj
0.83 0.64 0.02 0 k
(12)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(14)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
3.2088 0 Td
(20)Tj
0 g
(]\))Tj
1.8482 0 Td
(or)Tj
1.0998 0 Td
(die.)Tj
1.6611 0 Td
(The)Tj
1.7859 0 Td
(pro-)Tj
-35.1834 -1.3039 Td
(gression)Tj
3.5206 0 Td
(of)Tj
1.0148 0 Td
(visible)Tj
2.7495 0 Td
(signs)Tj
2.2224 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4683 0 Td
(disease)Tj
3.0161 0 Td
(can)Tj
1.627 0 Td
(be)Tj
1.1509 0 Td
(rapid,)Tj
2.5625 0 Td
(on)Tj
1.2756 0 Td
(the)Tj
1.4683 0 Td
(scale)Tj
2.109 0 Td
(of)Tj
1.0148 0 Td
(days.)Tj
2.2223 0 Td
(In)Tj
1.1112 0 Td
(fact,)Tj
1.8879 0 Td
(this)Tj
1.6724 0 Td
(short)Tj
2.2904 0 Td
(time-)Tj
-35.3988 -1.2983 Td
(frame)Tj
2.5568 0 Td
(has)Tj
1.542 0 Td
(led)Tj
1.4173 0 Td
(some)Tj
2.3301 0 Td
(pathologists)Tj
5.0116 0 Td
(to)Tj
1.0262 0 Td
(argue)Tj
2.4377 0 Td
(that)Tj
1.7802 0 Td
(wasting)Tj
4.0648 0 Td
(disease)Tj
3.0161 0 Td
(is)Tj
0.8447 0 Td
(a)Tj
0.6576 0 Td
(misnomer,)Tj
4.5638 0 Td
(as)Tj
1.0148 0 Td
(this)Tj
1.6724 0 Td
(term)Tj
-33.9361 -1.2983 Td
(suggests)Tj
3.4809 0 Td
(a)Tj
0.6519 0 Td
(gradual)Tj
3.2315 0 Td
(reduction)Tj
4.1102 0 Td
(in)Tj
1.0375 0 Td
(body)Tj
2.2167 0 Td
(mass)Tj
2.194 0 Td
(\(M.M.)Tj
2.7949 0 Td
(Garner)Tj
3.0841 0 Td
(pers.)Tj
2.1203 0 Td
(com.\).)Tj
2.7609 0 Td
(After)Tj
2.2847 0 Td
(SSWD-induced)Tj
-29.9676 -1.3039 Td
(mortality,)Tj
4.1385 0 Td
(decomposition)Tj
6.1965 0 Td
(occurs)Tj
2.8346 0 Td
(quickly.)Tj
3.3676 0 Td
(Although)Tj
4.0025 0 Td
(observers)Tj
4.0025 0 Td
(in)Tj
1.0374 0 Td
(low-flow)Tj
3.7304 0 Td
(subtidal)Tj
3.3902 0 Td
(systems)Tj
-32.7002 -1.2983 Td
(have)Tj
2.0579 0 Td
(noted)Tj
2.5285 0 Td
(piles)Tj
2.0296 0 Td
(of)Tj
1.0204 0 Td
(disarticulated)Tj
5.6239 0 Td
(ossicles)Tj
3.1692 0 Td
(\(skeletal)Tj
3.4922 0 Td
(elements\))Tj
4.1046 0 Td
(as)Tj
1.0148 0 Td
(indicators)Tj
4.2235 0 Td
(of)Tj
1.0148 0 Td
(recent)Tj
2.7099 0 Td
(SSWD-)Tj
ET
Q
q
1 j
1 J
0 w
576 737.1 m
36 737.1 l
36 737.6 l
576 737.6 l
f*
36 741.2598 107.1496 23.6976 re
W* n
q
107.0929 0 0 23.6409 36 741.3165 cm
q
/I0 Do
Q
Q
Q
q
0 0 612 792 re
W* n
1 j
1 J
0 w
7.9999 0 0 7.9999 397.0771 745.7952 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(Large-scale)Tj
5.3504 0 Td
(impacts)Tj
3.6071 0 Td
(of)Tj
1.0417 0 Td
(SSWD)Tj
3.1961 0 Td
(on)Tj
1.3111 0 Td
(interti)Tj
2.3882 0 Td
(dal)Tj
1.5236 0 Td
(sea)Tj
1.8071 0 Td
(stars)Tj
ET
Q
q
1 j
1 J
0 w
36 48.0002 m
576 48.0002 l
576 47.5002 l
36 47.5002 l
f*
0.83 0.64 0.02 0 k
81.5811 34.9228 m
237.8835 34.9228 l
h
f*
0 g
7.9999 0 0 7.9999 36 36 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(PLOS)Tj
2.8559 0 Td
(ONE)Tj
2.3669 0 Td
(|)Tj
0.83 0.64 0.02 0 k
0.4748 0 Td
(https://doi.or)Tj
5.4142 0 Td
(g/10.137)Tj
3.8198 0 Td
(1/journal.po)Tj
5.1236 0 Td
(ne.01928)Tj
4.089 0 Td
(70)Tj
0 g
2.0906 0 Td
(March)Tj
2.9622 0 Td
(20,)Tj
1.5874 0 Td
(2018)Tj
34.3562 0 Td
(2)Tj
0.7654 0 Td
(/)Tj
0.496 0 Td
(21)Tj
ET
Q
q
0.83 0.64 0.02 0 k
1 j
1 J
0 w
79.1433 685.6441 m
148.989 685.6441 l
h
f*
98.9858 663.6472 m
174.3874 663.6472 l
h
f*
38.4378 619.6535 m
102.2173 619.6535 l
h
f*
114.7465 608.6551 m
166.1669 608.6551 l
h
f*
0 g
7.9999 0 0 7.9999 36 708.7181 cm
BT
/F8 1 Tf
1 TL
-0.025 Tc
0 0 Td
(Funding:)Tj
/F9 1 Tf
3.5716 0 Td
(Research)Tj
3.6922 0 Td
(by)Tj
1.1197 0 Td
(MARINe)Tj
3.352 0 Td
(was)Tj
1.7008 0 Td
(primarily)Tj
-13.4363 -1.3748 Td
(sponsored)Tj
4.1598 0 Td
(by)Tj
1.1126 0 Td
(the)Tj
1.3748 0 Td
(Bureau)Tj
2.8418 0 Td
(of)Tj
0.9425 0 Td
(Ocean)Tj
2.5512 0 Td
(Energy)Tj
-12.9827 -1.3748 Td
(Manageme)Tj
4.1386 0 Td
(nt)Tj
0.9496 0 Td
(\()Tj
0.83 0.64 0.02 0 k
(https://www)Tj
4.8402 0 Td
(.boem.gov/)Tj
0 g
(,)Tj
4.6417 0 Td
(grant)Tj
2.1473 0 Td
(to)Tj
-16.7174 -1.3748 Td
(PTR)Tj
1.8071 0 Td
(#443634)Tj
3.5362 0 Td
(82772\),)Tj
3.1181 0 Td
(the)Tj
1.3678 0 Td
(David)Tj
2.3173 0 Td
(and)Tj
1.5874 0 Td
(Lucile)Tj
-13.7339 -1.3748 Td
(Packard)Tj
3.2102 0 Td
(Foundation)Tj
4.3512 0 Td
(\()Tj
0.83 0.64 0.02 0 k
(https://www)Tj
4.8402 0 Td
(.packard.org)Tj
4.6418 0 Td
(/)Tj
0 g
(,)Tj
-17.0434 -1.3748 Td
(grant)Tj
2.1543 0 Td
(to)Tj
0.9425 0 Td
(PTR)Tj
1.8071 0 Td
(#443634)Tj
3.5363 0 Td
(63620)Tj
2.5866 0 Td
(BIPRIT\),)Tj
3.3733 0 Td
(the)Tj
1.3677 0 Td
(National)Tj
-15.7678 -1.3748 Td
(Science)Tj
3.0968 0 Td
(Foundation)Tj
4.3512 0 Td
(\(grants)Tj
2.8843 0 Td
(to)Tj
0.9425 0 Td
(PTR)Tj
1.8071 0 Td
(#OCE-17373)Tj
4.7339 0 Td
(72)Tj
-17.8158 -1.3748 Td
(and)Tj
1.5874 0 Td
(OCE-17356)Tj
4.2591 0 Td
(07\),)Tj
1.7008 0 Td
(the)Tj
1.3677 0 Td
(U.S.)Tj
1.7858 0 Td
(Nationa)Tj
2.8205 0 Td
(l)Tj
0.4181 0 Td
(Park)Tj
1.8922 0 Td
(Service)Tj
-15.8316 -1.3749 Td
(\()Tj
0.83 0.64 0.02 0 k
(https://w)Tj
3.5575 0 Td
(ww.nps.gov)Tj
4.4646 0 Td
(/)Tj
0 g
(\),)Tj
1.0063 0 Td
(and)Tj
1.5945 0 Td
(the)Tj
1.3677 0 Td
(State)Tj
2.0976 0 Td
(of)Tj
0.9497 0 Td
(California)Tj
-15.0379 -1.3748 Td
(Ocean)Tj
2.5441 0 Td
(Protection)Tj
4.0181 0 Td
(Council)Tj
2.9764 0 Td
(\()Tj
0.83 0.64 0.02 0 k
(www.opc.)Tj
4.0394 0 Td
(ca.gov/)Tj
0 g
(,)Tj
3.1394 0 Td
(grant)Tj
-16.7174 -1.3748 Td
(to)Tj
0.9496 0 Td
(PTR)Tj
1.8071 0 Td
(#0CA15032\))Tj
4.6701 0 Td
(.)Tj
0.4464 0 Td
(BOEM)Tj
2.5583 0 Td
(scientists)Tj
3.6993 0 Td
(assisted)Tj
3.2457 0 Td
(with)Tj
-17.3765 -1.3748 Td
(data)Tj
1.7929 0 Td
(collection)Tj
3.7347 0 Td
(at)Tj
0.9 0 Td
(some)Tj
2.289 0 Td
(sites.)Tj
2.1614 0 Td
(Data)Tj
1.8992 0 Td
(from)Tj
2.0126 0 Td
(sites)Tj
1.9347 0 Td
(within)Tj
-16.7245 -1.3748 Td
(U.S.)Tj
1.7787 0 Td
(National)Tj
3.2386 0 Td
(Parks)Tj
2.3174 0 Td
(was)Tj
1.7078 0 Td
(primari)Tj
2.6434 0 Td
(ly)Tj
0.8362 0 Td
(collected)Tj
3.4937 0 Td
(by)Tj
-16.0158 -1.3748 Td
(National)Tj
3.2386 0 Td
(Park)Tj
1.8992 0 Td
(Service)Tj
2.9268 0 Td
(scientists,)Tj
3.926 0 Td
(who)Tj
1.8 0 Td
(also)Tj
1.7362 0 Td
(assisted)Tj
-15.5268 -1.3748 Td
(with)Tj
1.7787 0 Td
(manuscript)Tj
4.415 0 Td
(preparation.)Tj
4.6701 0 Td
(Support)Tj
3.1961 0 Td
(was)Tj
-14.0599 -1.3749 Td
(provided)Tj
3.4582 0 Td
(to)Tj
0.9497 0 Td
(CDGH)Tj
2.5086 0 Td
(by)Tj
1.1126 0 Td
(the)Tj
1.3678 0 Td
(Natural)Tj
2.8842 0 Td
(Sciences)Tj
3.515 0 Td
(and)Tj
-15.7961 -1.3748 Td
(Engineering)Tj
4.5992 0 Td
(Researc)Tj
2.9977 0 Td
(h)Tj
0.6945 0 Td
(Council)Tj
2.9764 0 Td
(of)Tj
0.9496 0 Td
(Canada)Tj
2.9551 0 Td
(Discovery)Tj
-15.1725 -1.3748 Td
(Grant)Tj
2.2677 0 Td
(\(RGPIN-)Tj
3.1536 0 Td
(2016-05441\))Tj
4.8756 0 Td
(.)Tj
0.4464 0 Td
(This)Tj
1.7788 0 Td
(is)Tj
0.8362 0 Td
(contributio)Tj
3.9898 0 Td
(n)Tj
-17.3481 -1.3819 Td
(number)Tj
3.1252 0 Td
(481)Tj
1.637 0 Td
(from)Tj
2.0126 0 Td
(PISCO,)Tj
2.863 0 Td
(the)Tj
1.3677 0 Td
(Partnership)Tj
4.493 0 Td
(for)Tj
-15.4985 -1.3748 Td
(Interdiscipli)Tj
4.2732 0 Td
(nary)Tj
1.8425 0 Td
(Studies)Tj
2.9835 0 Td
(of)Tj
0.9496 0 Td
(Coastal)Tj
2.9339 0 Td
(Oceans:)Tj
3.189 0 Td
(a)Tj
0.6449 0 Td
(long-)Tj
-16.8166 -1.3748 Td
(term)Tj
1.9559 0 Td
(Ecological)Tj
3.9614 0 Td
(Consorti)Tj
3.1182 0 Td
(um)Tj
1.4527 0 Td
(funded)Tj
2.7851 0 Td
(by)Tj
1.1197 0 Td
(the)Tj
1.3677 0 Td
(David)Tj
-15.7607 -1.3748 Td
(and)Tj
1.5874 0 Td
(Lucile)Tj
2.3953 0 Td
(Packard)Tj
3.2102 0 Td
(Foundation)Tj
4.3583 0 Td
(and)Tj
1.5874 0 Td
(the)Tj
1.3678 0 Td
(Gordon)Tj
3.0118 0 Td
(and)Tj
-17.5182 -1.3748 Td
(Betty)Tj
2.0976 0 Td
(Moore)Tj
2.6504 0 Td
(Foundation.)Tj
/F8 1 Tf
-4.748 -2.119 Td
(Competing)Tj
4.3795 0 Td
(interests)Tj
3.3307 0 Td
(:)Tj
/F9 1 Tf
0.4819 0 Td
(The)Tj
1.5875 0 Td
(authors)Tj
3.0401 0 Td
(have)Tj
1.963 0 Td
(declared)Tj
-14.7827 -1.3819 Td
(that)Tj
1.6228 0 Td
(no)Tj
1.1693 0 Td
(competing)Tj
4.1599 0 Td
(interests)Tj
ET
endstream
endobj
56 0 obj
<>stream
endstream
endobj
57 0 obj
<>stream
endstream
endobj
58 0 obj
<>stream
endstream
endobj
59 0 obj
<>stream
endstream
endobj
60 0 obj
<>stream
endstream
endobj
61 0 obj
<>stream
endstream
endobj
62 0 obj
<>stream
endstream
endobj
63 0 obj
<>stream
endstream
endobj
64 0 obj
<>stream
endstream
endobj
65 0 obj
<>stream
endstream
endobj
66 0 obj
<>stream
endstream
endobj
67 0 obj
<>stream
endstream
endobj
68 0 obj
<>stream
endstream
endobj
69 0 obj
<>stream
endstream
endobj
70 0 obj
<>stream
endstream
endobj
71 0 obj
<>stream
endstream
endobj
72 0 obj
<>stream
endstream
endobj
73 0 obj
<>stream
endstream
endobj
74 0 obj
<>stream
endstream
endobj
75 0 obj
<>stream
BT
10.3394 -33.7538 Td
(exist.)Tj
ET
Q
endstream
endobj
76 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 12 0 R/Annots 77 0 R/Contents 78 0 R/TrimBox[0 0 612 792]>>
endobj
77 0 obj
[79 0 R 80 0 R 81 0 R 82 0 R 83 0 R 84 0 R 85 0 R 86 0 R 87 0 R 88 0 R 89 0 R 90 0 R 91 0 R 92 0 R 93 0 R 94 0 R 95 0 R]
endobj
79 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref021)>>
endobj
80 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref022)>>
endobj
81 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref023)>>
endobj
82 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref024)>>
endobj
83 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref008)>>
endobj
84 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref025)>>
endobj
85 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref027)>>
endobj
86 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref004)>>
endobj
87 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref002)>>
endobj
88 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref004)>>
endobj
89 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref001)>>
endobj
90 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref002)>>
endobj
91 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref004)>>
endobj
92 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref027)>>
endobj
93 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref029)>>
endobj
94 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.ref030)>>
endobj
95 0 obj
<>/Border[0 0 0]/A 96 0 R>>
endobj
96 0 obj
<>
endobj
78 0 obj
[97 0 R 98 0 R 99 0 R 100 0 R 101 0 R 102 0 R 103 0 R 104 0 R 105 0 R 106 0 R 107 0 R 108 0 R 109 0 R]
endobj
97 0 obj
<>stream
q
0.83 0.64 0.02 0 k
294.0661 589.3228 m
303.5906 589.3228 l
h
f*
402.1228 589.3228 m
411.6472 589.3228 l
h
f*
511.0299 589.3228 m
520.5543 589.3228 l
h
f*
235.8425 537.3354 m
245.3669 537.3354 l
h
f*
401.1591 511.3134 m
405.9213 511.3134 l
h
f*
408.189 511.3134 m
417.7134 511.3134 l
h
f*
422.8724 511.3134 m
432.3969 511.3134 l
h
f*
225.1276 459.326 m
229.8898 459.326 l
h
f*
203.4142 420.3213 m
208.1764 420.3213 l
h
f*
210.3874 420.3213 m
215.1496 420.3213 l
h
f*
203.4142 407.3386 m
208.1764 407.3386 l
h
f*
210.3874 407.3386 m
215.1496 407.3386 l
h
f*
217.4173 407.3386 m
222.1795 407.3386 l
h
f*
224.4472 407.3386 m
233.9717 407.3386 l
h
f*
239.1307 407.3386 m
248.6551 407.3386 l
h
f*
203.4142 381.3165 m
212.9386 381.3165 l
h
f*
0 g
1 j
1 J
0 w
10 0 0 10 200.0125 707.4141 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(induced)Tj
3.4639 0 Td
(sea)Tj
1.44 0 Td
(star)Tj
1.6837 0 Td
(mortality,)Tj
4.1443 0 Td
(water)Tj
2.4264 0 Td
(motion)Tj
3.1691 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4683 0 Td
(intertidal)Tj
3.9005 0 Td
(zone)Tj
2.1146 0 Td
(quickly)Tj
3.1408 0 Td
(disperses)Tj
3.8324 0 Td
(decomposed)Tj
-31.8215 -1.3039 Td
(bodies)Tj
2.8119 0 Td
(of)Tj
1.0148 0 Td
(diseased)Tj
3.5433 0 Td
(asteroids)Tj
3.77 0 Td
(after)Tj
2.0353 0 Td
(death.)Tj
2.6589 0 Td
(Because)Tj
3.3959 0 Td
(of)Tj
1.0148 0 Td
(this)Tj
1.6724 0 Td
(combination)Tj
5.3461 0 Td
(of)Tj
1.0148 0 Td
(factors,)Tj
3.1294 0 Td
(monitoring)Tj
-31.4076 -1.2982 Td
(efforts)Tj
2.7552 0 Td
(initiated)Tj
3.5546 0 Td
(at)Tj
0.9525 0 Td
(the)Tj
1.4683 0 Td
(start)Tj
1.9842 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4684 0 Td
(disease)Tj
3.016 0 Td
(outbreak)Tj
3.7871 0 Td
(were)Tj
2.1203 0 Td
(unlikely)Tj
3.4299 0 Td
(to)Tj
1.0261 0 Td
(provide)Tj
3.2882 0 Td
(accurate)Tj
3.5546 0 Td
(character-)Tj
-33.4202 -1.2983 Td
(izations)Tj
3.3165 0 Td
(of)Tj
1.0148 0 Td
(changes)Tj
3.4015 0 Td
(in)Tj
/F10 1 Tf
1.0375 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0308 0 Td
(populations)Tj
4.9493 0 Td
(at)Tj
0.9524 0 Td
(any)Tj
1.6611 0 Td
(given)Tj
2.3755 0 Td
(location.)Tj
3.6793 0 Td
(Without)Tj
3.6113 0 Td
(pre-outbreak)Tj
-31.0221 -1.2983 Td
(data)Tj
1.9105 0 Td
(to)Tj
1.0261 0 Td
(provide)Tj
3.2882 0 Td
(historical)Tj
3.9231 0 Td
(context,)Tj
3.4129 0 Td
(it)Tj
0.7824 0 Td
(is)Tj
0.8504 0 Td
(impossible)Tj
4.4957 0 Td
(to)Tj
1.0261 0 Td
(evaluate)Tj
3.4639 0 Td
(the)Tj
1.4684 0 Td
(impact)Tj
2.9763 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.474 0 Td
(disease)Tj
3.0161 0 Td
(or)Tj
1.0998 0 Td
(the)Tj
-35.2287 -1.3039 Td
(trajectory)Tj
4.0591 0 Td
(of)Tj
1.0148 0 Td
(recovery)Tj
3.6454 0 Td
(for)Tj
1.3889 0 Td
(a)Tj
/F10 1 Tf
0.652 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9978 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0308 0 Td
(population.)Tj
4.8132 0 Td
(In)Tj
1.1112 0 Td
(these)Tj
2.2507 0 Td
(circumstances,)Tj
6.1285 0 Td
(data)Tj
1.9162 0 Td
(from)Tj
2.2053 0 Td
(ongoing)Tj
-34.2139 -1.2982 Td
(monitoring)Tj
4.8188 0 Td
(programs)Tj
4.0649 0 Td
(provide)Tj
3.2881 0 Td
(the)Tj
1.474 0 Td
(best)Tj
1.8085 0 Td
(means)Tj
2.8063 0 Td
(of)Tj
1.0148 0 Td
(assessing)Tj
3.7984 0 Td
(the)Tj
1.474 0 Td
(impact)Tj
2.9764 0 Td
(of)Tj
1.0148 0 Td
(this)Tj
1.6781 0 Td
(disease.)Tj
-29.0209 -1.2983 Td
(This)Tj
1.9899 0 Td
(is)Tj
0.8447 0 Td
(not)Tj
1.576 0 Td
(the)Tj
1.4684 0 Td
(first)Tj
1.8085 0 Td
(documented)Tj
5.244 0 Td
(disease)Tj
3.0217 0 Td
(outbreak)Tj
3.7814 0 Td
(in)Tj
1.0375 0 Td
(asteroids)Tj
3.7701 0 Td
(on)Tj
1.2756 0 Td
(the)Tj
1.4683 0 Td
(North)Tj
2.6645 0 Td
(American)Tj
-31.1468 -1.3039 Td
(Pacific)Tj
2.8799 0 Td
(coast;)Tj
2.4718 0 Td
(since)Tj
2.2394 0 Td
(the)Tj
1.474 0 Td
(1970s,)Tj
2.7042 0 Td
(outbreaks)Tj
4.1499 0 Td
(of)Tj
1.0148 0 Td
(\(uncharacterized\))Tj
7.3134 0 Td
(diseases)Tj
3.3788 0 Td
(have)Tj
2.058 0 Td
(been)Tj
2.1259 0 Td
(documented)Tj
-31.8101 -1.2983 Td
(in)Tj
1.0318 0 Td
(southern)Tj
3.787 0 Td
(California)Tj
4.2463 0 Td
([)Tj
0.83 0.64 0.02 0 k
(21)Tj
0 g
(],)Tj
2.0749 0 Td
(the)Tj
1.474 0 Td
(Gulf)Tj
1.9956 0 Td
(of)Tj
1.0148 0 Td
(California)Tj
4.2463 0 Td
([)Tj
0.83 0.64 0.02 0 k
(22)Tj
0 g
(],)Tj
2.0749 0 Td
(and)Tj
1.7348 0 Td
(British)Tj
2.8914 0 Td
(Columbia)Tj
4.1895 0 Td
([)Tj
0.83 0.64 0.02 0 k
(23)Tj
0 g
(].)Tj
2.075 0 Td
(Data)Tj
2.1259 0 Td
(from)Tj
-34.9622 -1.2983 Td
(these)Tj
2.2506 0 Td
(prior)Tj
2.2564 0 Td
(disease)Tj
3.0161 0 Td
(events)Tj
2.7325 0 Td
(indicate)Tj
3.4073 0 Td
(that)Tj
1.7801 0 Td
(most)Tj
2.2053 0 Td
(outbreaks)Tj
4.1443 0 Td
(were)Tj
2.1259 0 Td
(spatially)Tj
3.4753 0 Td
(and)Tj
1.7348 0 Td
(temporally)Tj
4.5354 0 Td
(limited.)Tj
-33.664 -1.3039 Td
(For)Tj
1.627 0 Td
(example,)Tj
3.7701 0 Td
(in)Tj
1.0375 0 Td
(1997)Tj
2.1203 0 Td
(diseased)Tj
3.5489 0 Td
(individuals)Tj
4.6488 0 Td
(were)Tj
2.1203 0 Td
(reported)Tj
3.6623 0 Td
(only)Tj
1.9729 0 Td
(as)Tj
1.0205 0 Td
(far)Tj
1.3096 0 Td
(south)Tj
2.4435 0 Td
(as)Tj
1.0148 0 Td
(Punta)Tj
2.5851 0 Td
(Banda,)Tj
2.9707 0 Td
(Baja)Tj
-35.8523 -1.2982 Td
(California)Tj
4.2462 0 Td
(and)Tj
1.7291 0 Td
(as)Tj
1.0148 0 Td
(far)Tj
1.3153 0 Td
(north)Tj
2.4718 0 Td
(as)Tj
1.0148 0 Td
(Punto)Tj
2.6589 0 Td
(Estero)Tj
2.7496 0 Td
(California,)Tj
4.473 0 Td
(spanning)Tj
3.9118 0 Td
(approximately)Tj
5.9754 0 Td
(4)Tj
0.6973 0 Td
(degrees)Tj
3.2145 0 Td
(of)Tj
-35.4725 -1.2983 Td
(latitude)Tj
3.2428 0 Td
([)Tj
0.83 0.64 0.02 0 k
(24)Tj
0 g
(].)Tj
2.0749 0 Td
(The)Tj
1.7858 0 Td
(current)Tj
3.1805 0 Td
(epidemic)Tj
3.8891 0 Td
(is)Tj
0.8504 0 Td
(strikingly)Tj
3.9968 0 Td
(different)Tj
3.6567 0 Td
(from)Tj
2.2053 0 Td
(previous)Tj
3.6453 0 Td
(events)Tj
2.7326 0 Td
(in)Tj
1.0375 0 Td
(both)Tj
2.0579 0 Td
(geo-)Tj
-34.3556 -1.3039 Td
(graphic)Tj
3.2258 0 Td
(extent)Tj
2.6758 0 Td
(and)Tj
1.7292 0 Td
(persistence,)Tj
4.8642 0 Td
(and)Tj
1.7291 0 Td
(has)Tj
1.5477 0 Td
(contributed)Tj
4.9153 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.474 0 Td
(growing)Tj
3.5263 0 Td
(push)Tj
2.1543 0 Td
(for)Tj
1.3833 0 Td
(the)Tj
1.474 0 Td
(development)Tj
-31.7251 -1.2983 Td
(of)Tj
1.0147 0 Td
(coordinated)Tj
5.057 0 Td
(responses)Tj
4.0932 0 Td
(to)Tj
1.0262 0 Td
(marine)Tj
3.084 0 Td
(diseases)Tj
3.3732 0 Td
(\(e.g.,)Tj
2.126 0 Td
([)Tj
0.83 0.64 0.02 0 k
(8)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(25)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(27)Tj
0 g
(]\).)Tj
4.5864 0 Td
(An)Tj
1.457 0 Td
(underlying)Tj
4.5978 0 Td
(goal)Tj
1.8709 0 Td
(of)Tj
1.0148 0 Td
(this)Tj
1.6781 0 Td
(coor-)Tj
-34.9793 -1.2982 Td
(dinated)Tj
3.2484 0 Td
(effort)Tj
2.3925 0 Td
(would)Tj
2.7155 0 Td
(be)Tj
1.1509 0 Td
(to)Tj
1.0205 0 Td
(develop)Tj
3.3221 0 Td
(better)Tj
2.5399 0 Td
(forecasting)Tj
4.6091 0 Td
(tools)Tj
2.1373 0 Td
(for)Tj
1.3889 0 Td
(disease)Tj
3.0161 0 Td
(events.)Tj
2.9537 0 Td
(With)Tj
2.279 0 Td
(a)Tj
0.6577 0 Td
(vast)Tj
1.7688 0 Td
(num-)Tj
-35.2004 -1.2983 Td
(ber)Tj
1.5193 0 Td
(of)Tj
1.0148 0 Td
(conditions)Tj
4.4787 0 Td
(and)Tj
1.7291 0 Td
(stressors)Tj
3.634 0 Td
(that)Tj
1.7802 0 Td
(could)Tj
2.4491 0 Td
(be)Tj
1.1509 0 Td
(associated)Tj
4.2576 0 Td
(with)Tj
1.9955 0 Td
(any)Tj
1.6611 0 Td
(disease)Tj
3.0161 0 Td
(outbreak,)Tj
4.0081 0 Td
(identifying)Tj
-32.6945 -1.3039 Td
(specific)Tj
3.1917 0 Td
(`warning)Tj
3.7644 0 Td
(signs')Tj
2.4378 0 Td
(and)Tj
1.7291 0 Td
(generating)Tj
4.4617 0 Td
(predictions)Tj
4.7339 0 Td
(for)Tj
1.3889 0 Td
(disease)Tj
3.0161 0 Td
(onset)Tj
2.3584 0 Td
(or)Tj
1.0998 0 Td
(impact)Tj
2.9764 0 Td
(are)Tj
1.4513 0 Td
(not)Tj
1.5704 0 Td
(simple)Tj
-34.1799 -1.2983 Td
(tasks)Tj
2.1713 0 Td
([)Tj
0.83 0.64 0.02 0 k
(4)Tj
0 g
(].)Tj
1.5987 0 Td
(Yet)Tj
1.5761 0 Td
(among)Tj
2.9933 0 Td
(these)Tj
2.2507 0 Td
(many)Tj
2.4775 0 Td
(possibilities,)Tj
5.0966 0 Td
(two)Tj
1.7122 0 Td
(factors)Tj
2.9026 0 Td
(stand)Tj
2.3868 0 Td
(out)Tj
1.5534 0 Td
(with)Tj
2.0012 0 Td
(strong)Tj
2.7666 0 Td
(potential)Tj
3.7644 0 Td
(for)Tj
-35.2514 -1.2983 Td
(predicting)Tj
4.3313 0 Td
(disease)Tj
3.016 0 Td
(affecting)Tj
3.6567 0 Td
(sea)Tj
1.4343 0 Td
(stars:)Tj
2.2734 0 Td
(temperature)Tj
5.1306 0 Td
(and)Tj
1.7292 0 Td
(population)Tj
4.5864 0 Td
(density.)Tj
3.3278 0 Td
(Both)Tj
2.1317 0 Td
(have)Tj
2.0636 0 Td
(been)Tj
-33.681 -1.3039 Td
(linked)Tj
2.7269 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.4683 0 Td
(emergence)Tj
4.5468 0 Td
(and)Tj
1.7291 0 Td
(severity)Tj
3.2712 0 Td
(of)Tj
1.0148 0 Td
(prior)Tj
2.2563 0 Td
(disease)Tj
3.0161 0 Td
(events)Tj
2.7325 0 Td
(in)Tj
1.0319 0 Td
(marine)Tj
3.084 0 Td
(systems)Tj
3.2995 0 Td
(\(reviewed)Tj
4.1386 0 Td
(in)Tj
-35.3421 -1.2982 Td
([)Tj
0.83 0.64 0.02 0 k
(2)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(4)Tj
0 g
(]\))Tj
2.4151 0 Td
(and)Tj
1.7291 0 Td
(are)Tj
1.4513 0 Td
(widely)Tj
2.8233 0 Td
(considered)Tj
4.5978 0 Td
(to)Tj
1.0261 0 Td
(be)Tj
1.1509 0 Td
(important)Tj
4.2802 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4684 0 Td
(development)Tj
5.4141 0 Td
(of)Tj
1.0148 0 Td
(predictive)Tj
4.1896 0 Td
(models)Tj
-32.5982 -1.2983 Td
([)Tj
0.83 0.64 0.02 0 k
(1)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(2)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(4)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
(27)Tj
0 g
()Tj
0.83 0.64 0.02 0 k
(29)Tj
0 g
(].)Tj
1.1962 -1.3039 Td
(Once)Tj
2.364 0 Td
(a)Tj
0.652 0 Td
(disease)Tj
3.0217 0 Td
(outbreak)Tj
3.7814 0 Td
(does)Tj
2.0409 0 Td
(occur,)Tj
2.6986 0 Td
(especially)Tj
4.0195 0 Td
(one)Tj
1.6951 0 Td
(associated)Tj
4.2576 0 Td
(with)Tj
1.9956 0 Td
(a)Tj
0.6576 0 Td
(mass)Tj
2.194 0 Td
(mortality)Tj
3.9175 0 Td
(event)Tj
-34.4917 -1.2983 Td
([)Tj
0.83 0.64 0.02 0 k
(30)Tj
0 g
(],)Tj
2.0749 0 Td
(the)Tj
1.4683 0 Td
(ability)Tj
2.6986 0 Td
(to)Tj
1.0205 0 Td
(assess)Tj
2.5228 0 Td
(the)Tj
1.4683 0 Td
(degree)Tj
2.8573 0 Td
(of)Tj
1.0148 0 Td
(population)Tj
4.5921 0 Td
(recovery)Tj
3.6454 0 Td
(becomes)Tj
3.685 0 Td
(critically)Tj
3.6566 0 Td
(important.)Tj
4.5071 0 Td
(His-)Tj
-35.2117 -1.2983 Td
(torical)Tj
2.7665 0 Td
(data)Tj
1.9163 0 Td
(provided)Tj
3.8154 0 Td
(by)Tj
1.1792 0 Td
(long-term)Tj
4.2519 0 Td
(population)Tj
4.5921 0 Td
(surveillance)Tj
4.9436 0 Td
(are)Tj
1.4513 0 Td
(essential)Tj
3.583 0 Td
(at)Tj
0.9524 0 Td
(this)Tj
1.6725 0 Td
(stage,)Tj
2.4264 0 Td
(as)Tj
1.0148 0 Td
(they)Tj
-34.5654 -1.2982 Td
(allow)Tj
2.3357 0 Td
(the)Tj
1.4683 0 Td
(construction)Tj
5.3008 0 Td
(of)Tj
1.0204 0 Td
(`benchmark')Tj
5.2101 0 Td
(indices)Tj
3.0387 0 Td
(based)Tj
2.4718 0 Td
(on)Tj
1.2756 0 Td
(normal)Tj
3.9288 0 Td
(pre-disease)Tj
4.6828 0 Td
(population)Tj
-30.733 -1.3039 Td
(parameters,)Tj
4.8982 0 Td
(such)Tj
2.0579 0 Td
(as)Tj
1.0148 0 Td
(size)Tj
1.6838 0 Td
(frequency)Tj
4.1896 0 Td
(distributions)Tj
5.3291 0 Td
(and)Tj
1.7291 0 Td
(abundance.)Tj
4.8245 0 Td
(A)Tj
0.9014 0 Td
(variety)Tj
2.9254 0 Td
(of)Tj
1.0148 0 Td
(components)Tj
-30.5686 -1.2983 Td
(factor)Tj
2.5398 0 Td
(into)Tj
1.8425 0 Td
(population)Tj
4.5921 0 Td
(recovery,)Tj
3.8721 0 Td
(including)Tj
4.0422 0 Td
(recruitment)Tj
4.9946 0 Td
(of)Tj
1.0148 0 Td
(new)Tj
1.8765 0 Td
(individuals,)Tj
4.8699 0 Td
(juvenile)Tj
3.3732 0 Td
(survivor-)Tj
-33.0177 -1.2983 Td
(ship)Tj
1.8935 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.4684 0 Td
(adult)Tj
2.2506 0 Td
(stage,)Tj
2.4322 0 Td
(and)Tj
1.7291 0 Td
(disease)Tj
3.016 0 Td
(persistence.)Tj
4.8642 0 Td
(Each)Tj
2.177 0 Td
(of)Tj
1.0148 0 Td
(these)Tj
2.2507 0 Td
(components)Tj
5.1761 0 Td
(can)Tj
1.627 0 Td
(vary)Tj
1.9333 0 Td
(among)Tj
-32.859 -1.3039 Td
(affected)Tj
3.3221 0 Td
(populations,)Tj
5.1704 0 Td
(thus)Tj
1.9275 0 Td
(it)Tj
0.7767 0 Td
(is)Tj
0.8447 0 Td
(necessary)Tj
4.0139 0 Td
(to)Tj
1.0204 0 Td
(have)Tj
2.0523 0 Td
(long-term)Tj
4.2463 0 Td
(coordinated)Tj
5.0513 0 Td
(monitoring)Tj
4.8132 0 Td
(of)Tj
1.0091 0 Td
(multiple)Tj
-34.2479 -1.2982 Td
(populations)Tj
4.9492 0 Td
(in)Tj
1.0375 0 Td
(order)Tj
2.4208 0 Td
(to:)Tj
1.2472 0 Td
(a\))Tj
0.9978 0 Td
(build)Tj
2.296 0 Td
(recovery)Tj
3.6454 0 Td
(benchmark)Tj
4.7735 0 Td
(targets)Tj
2.8743 0 Td
(based)Tj
2.4718 0 Td
(on)Tj
1.2756 0 Td
(specific)Tj
3.1918 0 Td
(data,)Tj
2.1373 0 Td
(b\))Tj
1.0658 0 Td
(make)Tj
-34.384 -1.2983 Td
(informed)Tj
3.9741 0 Td
(assessments)Tj
4.9719 0 Td
(of)Tj
1.0148 0 Td
(recovery)Tj
3.6454 0 Td
(at)Tj
0.9524 0 Td
(various)Tj
3.1464 0 Td
(scales)Tj
2.4662 0 Td
(\(e.g.)Tj
1.8992 0 Td
(within)Tj
2.8119 0 Td
(and)Tj
1.7348 0 Td
(among)Tj
2.9934 0 Td
(regions\),)Tj
3.7303 0 Td
(and)Tj
1.7292 0 Td
(c\))Tj
-35.07 -1.3039 Td
(compare)Tj
3.7133 0 Td
(recovery)Tj
3.651 0 Td
(estimates)Tj
3.9061 0 Td
(at)Tj
0.9525 0 Td
(various)Tj
3.1407 0 Td
(scales)Tj
2.4718 0 Td
(in)Tj
1.0375 0 Td
(order)Tj
2.4208 0 Td
(to)Tj
1.0204 0 Td
(identify)Tj
3.2995 0 Td
(factors)Tj
2.9027 0 Td
(that)Tj
1.7745 0 Td
(might)Tj
2.6022 0 Td
(facilitate)Tj
3.5999 0 Td
(or)Tj
-36.4929 -1.2983 Td
(hinder)Tj
2.8856 0 Td
(recovery.)Tj
-1.6894 -1.2983 Td
(Here)Tj
2.211 0 Td
(we)Tj
1.3266 0 Td
(have)Tj
2.0579 0 Td
(combined)Tj
4.2463 0 Td
(data)Tj
1.9105 0 Td
(from)Tj
2.2054 0 Td
(several)Tj
2.9366 0 Td
(long-term)Tj
4.252 0 Td
(monitoring)Tj
4.8189 0 Td
(programs)Tj
4.0648 0 Td
(from)Tj
2.2054 0 Td
(mainland)Tj
-33.4316 -1.3039 Td
(and)Tj
1.7291 0 Td
(island)Tj
2.6022 0 Td
(sites)Tj
1.9332 0 Td
(along)Tj
2.4207 0 Td
(the)Tj
1.4684 0 Td
(North)Tj
2.6645 0 Td
(American)Tj
4.1896 0 Td
(Pacific)Tj
2.88 0 Td
(coast,)Tj
2.4718 0 Td
(ranging)Tj
3.3165 0 Td
(from)Tj
2.2053 0 Td
(Sitka,)Tj
2.4208 0 Td
(Alaska)Tj
2.8743 0 Td
(to)Tj
1.0262 0 Td
(San)Tj
-34.2026 -1.2982 Td
(Diego,)Tj
2.8402 0 Td
(California.)Tj
4.4731 0 Td
(Most)Tj
2.279 0 Td
(of)Tj
1.0148 0 Td
(these)Tj
2.2564 0 Td
(surveys)Tj
3.1748 0 Td
(were)Tj
2.1203 0 Td
(completed)Tj
4.405 0 Td
(by)Tj
1.1849 0 Td
(partners)Tj
3.5432 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4684 0 Td
(Multi-Agency)Tj
-29.7976 -1.2983 Td
(Rocky)Tj
2.7212 0 Td
(Intertidal)Tj
3.9741 0 Td
(Network)Tj
3.7361 0 Td
(\(MARINe\),)Tj
4.8245 0 Td
(who)Tj
1.9389 0 Td
(have)Tj
2.0636 0 Td
(been)Tj
2.1203 0 Td
(working)Tj
3.5603 0 Td
(with)Tj
1.9956 0 Td
(a)Tj
0.6576 0 Td
(standardized)Tj
5.3518 0 Td
(set)Tj
1.3039 0 Td
(of)Tj
1.0148 0 Td
(mon-)Tj
-35.2627 -1.2983 Td
(itoring)Tj
2.9423 0 Td
(protocols)Tj
3.9571 0 Td
(to)Tj
1.0262 0 Td
(study)Tj
2.3924 0 Td
(community)Tj
4.8869 0 Td
(dynamics)Tj
4.0535 0 Td
(in)Tj
1.0375 0 Td
(rocky)Tj
2.4718 0 Td
(intertidal)Tj
3.9004 0 Td
(systems)Tj
3.2995 0 Td
(for)Tj
1.389 0 Td
(up)Tj
1.2642 0 Td
(to)Tj
1.0262 0 Td
(25)Tj
1.1678 0 Td
(years.)Tj
-34.8148 -1.3039 Td
(This)Tj
1.9842 0 Td
(unique)Tj
3.0217 0 Td
(data)Tj
1.9162 0 Td
(set)Tj
1.3039 0 Td
(provides)Tj
3.6454 0 Td
(the)Tj
1.474 0 Td
(necessary)Tj
4.0195 0 Td
(temporal)Tj
3.8381 0 Td
(context)Tj
3.1804 0 Td
(to)Tj
1.0262 0 Td
(allow)Tj
2.3414 0 Td
(us)Tj
1.1055 0 Td
(to)Tj
1.0261 0 Td
(assess)Tj
/F10 1 Tf
2.5228 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
-33.3975 -1.2983 Td
(population)Tj
4.5864 0 Td
(changes,)Tj
3.634 0 Td
(and)Tj
1.7291 0 Td
(expansive)Tj
4.1159 0 Td
(spatial)Tj
2.7779 0 Td
(coverage)Tj
3.719 0 Td
(to)Tj
1.0262 0 Td
(allow)Tj
2.3357 0 Td
(us)Tj
1.1112 0 Td
(to)Tj
1.0261 0 Td
(evaluate)Tj
3.4583 0 Td
(the)Tj
1.474 0 Td
(impact)Tj
2.9763 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
-34.9849 -1.2982 Td
(disease)Tj
3.016 0 Td
(across)Tj
2.6759 0 Td
(nearly)Tj
2.6985 0 Td
(the)Tj
1.4741 0 Td
(entire)Tj
2.5511 0 Td
(species')Tj
3.2088 0 Td
(range.)Tj
2.6872 0 Td
(We)Tj
1.6101 0 Td
(use)Tj
1.5307 0 Td
(these)Tj
2.2564 0 Td
(data)Tj
1.9162 0 Td
(to:)Tj
1.2472 0 Td
(1\))Tj
1.0375 0 Td
(present)Tj
3.1634 0 Td
(regional)Tj
3.481 0 Td
(pat-)Tj
-34.5541 -1.3039 Td
(terns)Tj
2.2223 0 Td
(of)Tj
1.0148 0 Td
(difference)Tj
4.2066 0 Td
(in)Tj
1.0318 0 Td
(the)Tj
1.4683 0 Td
(degree)Tj
2.8573 0 Td
(to)Tj
1.0262 0 Td
(which)Tj
2.6475 0 Td
(SSWD)Tj
2.8743 0 Td
(has)Tj
1.5421 0 Td
(impacted)Tj
/F10 1 Tf
3.9344 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0309 0 Td
(populations,)Tj
5.176 0 Td
(2\))Tj
-35.0246 -1.2983 Td
(examine)Tj
3.5999 0 Td
(whether)Tj
3.4753 0 Td
(sea)Tj
1.44 0 Td
(star)Tj
1.6837 0 Td
(density)Tj
3.1011 0 Td
(might)Tj
2.5965 0 Td
(have)Tj
2.0637 0 Td
(played)Tj
2.8232 0 Td
(a)Tj
0.652 0 Td
(role)Tj
1.7688 0 Td
(in)Tj
1.0375 0 Td
(disease)Tj
3.016 0 Td
(severity,)Tj
3.498 0 Td
(3\))Tj
1.0374 0 Td
(explore)Tj
3.1748 0 Td
(the)Tj
-34.9679 -1.2983 Td
(potential)Tj
3.7587 0 Td
(relationship)Tj
4.9889 0 Td
(between)Tj
/F10 1 Tf
3.5263 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9978 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0308 0 Td
(decline)Tj
3.0784 0 Td
(and)Tj
1.7291 0 Td
(water)Tj
2.4322 0 Td
(temperature,)Tj
5.3574 0 Td
(which)Tj
2.6475 0 Td
(has)Tj
1.5421 0 Td
(been)Tj
-34.0892 -1.3039 Td
(implicated)Tj
4.4446 0 Td
(as)Tj
1.0148 0 Td
(a)Tj
0.652 0 Td
(contributing)Tj
5.2497 0 Td
(factor)Tj
2.5399 0 Td
(in)Tj
1.0374 0 Td
(prior)Tj
2.2564 0 Td
(wasting)Tj
3.2768 0 Td
(events,)Tj
2.9537 0 Td
(and)Tj
1.7291 0 Td
(4\))Tj
1.0375 0 Td
(present)Tj
3.1635 0 Td
(spatial)Tj
2.7779 0 Td
(patterns)Tj
3.4752 0 Td
(in)Tj
/F10 1 Tf
1.0318 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
-36.6403 -1.2982 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0308 0 Td
(recruitment)Tj
4.9946 0 Td
(and)Tj
1.7291 0 Td
(juvenile)Tj
3.3676 0 Td
(survival)Tj
3.3335 0 Td
(between)Tj
3.5319 0 Td
(pre-and)Tj
3.3902 0 Td
(post-)Tj
2.2564 0 Td
(outbreak)Tj
3.7871 0 Td
(periods)Tj
3.1918 0 Td
(that)Tj
1.7801 0 Td
(can)Tj
ET
Q
q
1 j
1 J
0 w
576 737.1 m
36 737.1 l
36 737.6 l
576 737.6 l
f*
36 741.2598 107.1496 23.6976 re
W* n
q
107.0929 0 0 23.6409 36 741.3165 cm
q
/I0 Do
Q
Q
Q
q
0 0 612 792 re
W* n
1 j
1 J
0 w
7.9999 0 0 7.9999 397.0771 745.7952 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(Large-scale)Tj
5.3504 0 Td
(impacts)Tj
3.6071 0 Td
(of)Tj
1.0417 0 Td
(SSWD)Tj
3.1961 0 Td
(on)Tj
1.3111 0 Td
(interti)Tj
2.3882 0 Td
(dal)Tj
1.5236 0 Td
(sea)Tj
1.8071 0 Td
(stars)Tj
ET
Q
q
1 j
1 J
0 w
36 48.0002 m
576 48.0002 l
576 47.5002 l
36 47.5002 l
f*
0.83 0.64 0.02 0 k
81.5811 34.9228 m
237.8835 34.9228 l
h
f*
0 g
7.9999 0 0 7.9999 36 36 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(PLOS)Tj
2.8559 0 Td
(ONE)Tj
2.3669 0 Td
(|)Tj
0.83 0.64 0.02 0 k
0.4748 0 Td
(https://doi.or)Tj
5.4142 0 Td
(g/10.137)Tj
3.8198 0 Td
(1/journal.po)Tj
5.1236 0 Td
(ne.01928)Tj
4.089 0 Td
(70)Tj
0 g
2.0906 0 Td
(March)Tj
2.9622 0 Td
(20,)Tj
1.5874 0 Td
(2018)Tj
34.3562 0 Td
(3)Tj
0.7654 0 Td
(/)Tj
ET
endstream
endobj
98 0 obj
<>stream
endstream
endobj
99 0 obj
<>stream
endstream
endobj
100 0 obj
<>stream
endstream
endobj
101 0 obj
<>stream
endstream
endobj
102 0 obj
<>stream
endstream
endobj
103 0 obj
<>stream
endstream
endobj
104 0 obj
<>stream
endstream
endobj
105 0 obj
<>stream
endstream
endobj
106 0 obj
<>stream
endstream
endobj
107 0 obj
<>stream
endstream
endobj
108 0 obj
<>stream
endstream
endobj
109 0 obj
<>stream
BT
66.402 0 Td
(21)Tj
ET
Q
endstream
endobj
110 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 12 0 R/Annots 112 0 R/Contents 113 0 R/TrimBox[0 0 612 792]>>
endobj
112 0 obj
[114 0 R 115 0 R 116 0 R 117 0 R 118 0 R]
endobj
114 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.g001)>>
endobj
115 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.s001)>>
endobj
116 0 obj
<>/Border[0 0 0]/Dest(Rpone.0192870.s003)>>
endobj
117 0 obj
<>/Border[0 0 0]/A 119 0 R>>
endobj
119 0 obj
<>
endobj
118 0 obj
<>/Border[0 0 0]/A 120 0 R>>
endobj
120 0 obj
<>
endobj
113 0 obj
[121 0 R 122 0 R 123 0 R 124 0 R 125 0 R 126 0 R 127 0 R 128 0 R]
endobj
121 0 obj
<>stream
q
0.83 0.64 0.02 0 k
245.1969 193.0394 m
264.5291 193.0394 l
h
f*
268.8945 193.0394 m
321.8457 193.0394 l
h
f*
326.211 193.0394 m
359.7165 193.0394 l
h
f*
419.9811 141.052 m
519.7039 141.052 l
h
f*
0 g
1 j
1 J
0 w
10 0 0 10 200.0125 707.4141 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(be)Tj
1.1508 0 Td
(used)Tj
2.0579 0 Td
(to)Tj
1.0262 0 Td
(5\))Tj
1.0375 0 Td
(assess)Tj
2.5171 0 Td
(the)Tj
1.474 0 Td
(potential)Tj
3.7587 0 Td
(for)Tj
1.3833 0 Td
(recovery)Tj
3.651 0 Td
(of)Tj
/F10 1 Tf
1.0148 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0309 0 Td
(populations)Tj
4.9549 0 Td
(within)Tj
2.812 0 Td
(SSWD-)Tj
-31.8612 -1.3039 Td
(impacted)Tj
3.9287 0 Td
(areas.)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 665.2913 cm
BT
/F2 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Materials)Tj
4.5071 0 Td
(and)Tj
1.989 0 Td
(methods)Tj
/F11 1 Tf
-6.4961 -1.5023 Td
(Ethics)Tj
2.7779 0 Td
(statement)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 630.2551 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(No)Tj
1.4626 0 Td
(animals)Tj
3.3279 0 Td
(were)Tj
2.1259 0 Td
(collected)Tj
3.7361 0 Td
(for)Tj
1.3889 0 Td
(these)Tj
2.2507 0 Td
(surveys,)Tj
3.4016 0 Td
(but)Tj
1.5534 0 Td
(many)Tj
2.4774 0 Td
(sites)Tj
1.9276 0 Td
(required)Tj
3.6396 0 Td
(permits)Tj
3.2825 0 Td
(for)Tj
1.3833 0 Td
(general)Tj
-31.9575 -1.2982 Td
(research)Tj
3.5546 0 Td
(or)Tj
1.0998 0 Td
(permission)Tj
4.6601 0 Td
(for)Tj
1.389 0 Td
(access.)Tj
2.8686 0 Td
(In)Tj
1.1112 0 Td
(Washington,)Tj
5.3688 0 Td
(work)Tj
2.279 0 Td
(at)Tj
0.9525 0 Td
(Post)Tj
1.9445 0 Td
(Point)Tj
2.3981 0 Td
(was)Tj
1.7008 0 Td
(conducted)Tj
4.4277 0 Td
(under)Tj
-33.7547 -1.2983 Td
(Washington)Tj
5.142 0 Td
(Department)Tj
5.091 0 Td
(of)Tj
1.0148 0 Td
(Fish)Tj
1.9048 0 Td
(and)Tj
1.7291 0 Td
(Wildlife)Tj
3.4526 0 Td
(\(DFW\))Tj
3.1295 0 Td
(permit)Tj
2.9139 0 Td
(#)Tj
0.6974 0 Td
(1207201)Tj
4.0648 0 Td
(and)Tj
1.7348 0 Td
(access)Tj
2.6419 0 Td
(to)Tj
1.0261 0 Td
(sites)Tj
1.9332 0 Td
(on)Tj
-36.4759 -1.3039 Td
(Makah)Tj
3.0047 0 Td
(land)Tj
1.9729 0 Td
(and)Tj
1.7347 0 Td
(the)Tj
1.4684 0 Td
(Quinault)Tj
3.8154 0 Td
(Nation)Tj
3.0104 0 Td
(was)Tj
1.7007 0 Td
(granted)Tj
3.2882 0 Td
(to)Tj
1.0261 0 Td
(Olympic)Tj
3.6964 0 Td
(Coast)Tj
2.4831 0 Td
(National)Tj
3.6907 0 Td
(Marine)Tj
3.1578 0 Td
(Sanctu-)Tj
-34.0495 -1.2983 Td
(ary)Tj
1.4796 0 Td
(\(NMS\))Tj
3.0047 0 Td
(for)Tj
1.3833 0 Td
(all)Tj
1.1509 0 Td
(surveys.)Tj
3.3959 0 Td
(In)Tj
1.1111 0 Td
(Oregon,)Tj
3.5036 0 Td
(surveys)Tj
3.1748 0 Td
(were)Tj
2.126 0 Td
(completed)Tj
4.405 0 Td
(under)Tj
2.6192 0 Td
(Oregon)Tj
3.2825 0 Td
(DFW)Tj
2.4491 0 Td
(permit)Tj
2.9197 0 Td
(#'s)Tj
-36.0054 -1.2982 Td
(18084,)Tj
2.8232 0 Td
(18610,)Tj
2.8177 0 Td
(19306,)Tj
2.8232 0 Td
(20174,)Tj
2.8233 0 Td
(and)Tj
1.7292 0 Td
(21411,)Tj
2.8233 0 Td
(and)Tj
1.7291 0 Td
(access)Tj
2.6418 0 Td
(to)Tj
1.0262 0 Td
(Fogarty)Tj
3.2768 0 Td
(Creek)Tj
2.5965 0 Td
(was)Tj
1.7008 0 Td
(provided)Tj
3.8154 0 Td
(through)Tj
3.4413 0 Td
(col-)Tj
-36.0678 -1.3039 Td
(laborators)Tj
4.2576 0 Td
(at)Tj
0.9524 0 Td
(Oregon)Tj
3.2825 0 Td
(State)Tj
2.1486 0 Td
(University,)Tj
4.6262 0 Td
(who)Tj
1.9388 0 Td
(have)Tj
2.058 0 Td
(an)Tj
1.2075 0 Td
(agreement)Tj
4.4277 0 Td
(with)Tj
1.9956 0 Td
(the)Tj
1.4683 0 Td
(owners.)Tj
3.3392 0 Td
(In)Tj
1.1112 0 Td
(California,)Tj
-32.8136 -1.2983 Td
(the)Tj
1.4683 0 Td
(National)Tj
3.6907 0 Td
(Oceanic)Tj
3.4866 0 Td
(and)Tj
1.7291 0 Td
(Atmospheric)Tj
5.4198 0 Td
(Administration)Tj
6.4233 0 Td
(Office)Tj
2.6532 0 Td
(of)Tj
1.0148 0 Td
(National)Tj
3.6963 0 Td
(Marine)Tj
3.1521 0 Td
(Sanctuaries)Tj
-32.7342 -1.2983 Td
(Program)Tj
3.7473 0 Td
(permitted)Tj
4.1669 0 Td
(surveys)Tj
3.1748 0 Td
(at)Tj
0.9524 0 Td
(sites)Tj
1.9276 0 Td
(within)Tj
2.8119 0 Td
(the)Tj
1.4741 0 Td
(Monterey)Tj
4.1385 0 Td
(Bay)Tj
1.6894 0 Td
(and)Tj
1.7292 0 Td
(Gulf)Tj
1.9955 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4684 0 Td
(Farallones)Tj
4.32 0 Td
(NMS)Tj
-34.6108 -1.3039 Td
(\(MBNMS-2010-001-A1,)Tj
9.9041 0 Td
(MBNMS-2015-015-A1,)Tj
9.5584 0 Td
(and)Tj
1.7348 0 Td
(MULTI-2017-010-A1\).)Tj
9.445 0 Td
(Work)Tj
2.5568 0 Td
(at)Tj
0.9524 0 Td
(other)Tj
-34.1515 -1.2983 Td
(California)Tj
4.2462 0 Td
(sites)Tj
1.9276 0 Td
(\(including)Tj
4.3823 0 Td
(Marine)Tj
3.1578 0 Td
(Protected)Tj
4.0535 0 Td
(Areas\))Tj
2.8346 0 Td
(was)Tj
1.7008 0 Td
(authorized)Tj
4.5127 0 Td
(by)Tj
1.1849 0 Td
(California)Tj
4.2463 0 Td
(DFW)Tj
2.4491 0 Td
(per-)Tj
-34.6958 -1.2982 Td
(mits)Tj
1.9672 0 Td
(SC-4055,)Tj
3.8324 0 Td
(SC-3124,)Tj
3.8381 0 Td
(SC-8187,)Tj
3.8381 0 Td
(SC-10589,)Tj
4.3143 0 Td
(and)Tj
1.7291 0 Td
(SC-003922.)Tj
4.7905 0 Td
(Annual)Tj
3.2088 0 Td
(permits)Tj
3.2768 0 Td
(to)Tj
1.0262 0 Td
(work)Tj
2.279 0 Td
(at)Tj
0.9525 0 Td
(sites)Tj
-35.053 -1.2983 Td
(located)Tj
3.0784 0 Td
(in)Tj
1.0374 0 Td
(California)Tj
4.2463 0 Td
(State)Tj
2.1486 0 Td
(Parks)Tj
2.4435 0 Td
(were)Tj
2.1203 0 Td
(granted)Tj
3.2882 0 Td
(by)Tj
1.1848 0 Td
(the)Tj
1.4684 0 Td
(California)Tj
4.2463 0 Td
(Department)Tj
5.0909 0 Td
(of)Tj
1.0148 0 Td
(Parks)Tj
2.4435 0 Td
(and)Tj
-33.8114 -1.3039 Td
(Recreation)Tj
4.5354 0 Td
(as)Tj
1.0148 0 Td
(follows:)Tj
3.2881 0 Td
(Channel)Tj
3.6113 0 Td
(Coast)Tj
2.4832 0 Td
(District)Tj
3.2428 0 Td
(State)Tj
2.1486 0 Td
(Parks)Tj
2.4435 0 Td
(to)Tj
1.0261 0 Td
(R.)Tj
1.0658 0 Td
(Ambrose)Tj
3.8948 0 Td
(and)Tj
1.7291 0 Td
(S.)Tj
0.9185 0 Td
(Lee,)Tj
1.8311 0 Td
(Crystal)Tj
-33.2331 -1.2983 Td
(Cove)Tj
2.2677 0 Td
(State)Tj
2.1486 0 Td
(Park)Tj
2.0806 0 Td
(to)Tj
1.0262 0 Td
(S.)Tj
0.9184 0 Td
(Murray,)Tj
3.4979 0 Td
(J.)Tj
0.7767 0 Td
(Smith,)Tj
2.8346 0 Td
(and)Tj
1.7291 0 Td
(J.)Tj
0.7767 0 Td
(Burnaford,)Tj
4.5978 0 Td
(and)Tj
1.7348 0 Td
(State)Tj
2.1486 0 Td
(Park)Tj
2.0807 0 Td
(system-wide)Tj
5.1873 0 Td
(permits)Tj
-33.8057 -1.2982 Td
(to)Tj
1.0261 0 Td
(P.)Tj
1.0034 0 Td
(Raimondi.)Tj
4.4391 0 Td
(Permission)Tj
4.7055 0 Td
(to)Tj
1.0204 0 Td
(work)Tj
2.2791 0 Td
(at)Tj
0.9524 0 Td
(Bodega,)Tj
3.3845 0 Td
(Coal)Tj
2.0693 0 Td
(Oil)Tj
1.4797 0 Td
(Point,)Tj
2.6249 0 Td
(Scripps)Tj
3.1521 0 Td
(Reef,)Tj
2.2053 0 Td
(and)Tj
1.7291 0 Td
(all)Tj
1.1452 0 Td
(sites)Tj
1.9332 0 Td
(on)Tj
-35.1493 -1.3039 Td
(Santa)Tj
2.4094 0 Td
(Cruz)Tj
2.194 0 Td
(Island)Tj
2.6815 0 Td
(was)Tj
1.6952 0 Td
(granted)Tj
3.2881 0 Td
(by)Tj
1.1849 0 Td
(the)Tj
1.4683 0 Td
(UC)Tj
1.6214 0 Td
(Natural)Tj
3.2655 0 Td
(Reserve)Tj
3.2995 0 Td
(System)Tj
3.0501 0 Td
(and)Tj
1.7348 0 Td
(the)Tj
1.4683 0 Td
(Nature)Tj
3.0104 0 Td
(Conser-)Tj
-32.3714 -1.2983 Td
(vancy.)Tj
2.7609 0 Td
(The)Tj
1.7801 0 Td
(Sea)Tj
1.5534 0 Td
(Ranch)Tj
2.7779 0 Td
(Association)Tj
4.8756 0 Td
(allowed)Tj
3.2882 0 Td
(access)Tj
2.6475 0 Td
(to)Tj
1.0261 0 Td
(our)Tj
1.6271 0 Td
(site)Tj
1.5704 0 Td
(there.)Tj
2.4888 0 Td
(An)Tj
1.4513 0 Td
(access)Tj
2.6476 0 Td
(agreement)Tj
4.4277 0 Td
(was)Tj
-34.9226 -1.2983 Td
(granted)Tj
3.2881 0 Td
(annually)Tj
3.6567 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.474 0 Td
(PISCO)Tj
3.0047 0 Td
(group)Tj
2.6079 0 Td
(at)Tj
0.9524 0 Td
(UCSC)Tj
2.7553 0 Td
(from)Tj
2.2053 0 Td
(the)Tj
1.474 0 Td
(Pebble)Tj
2.8857 0 Td
(Beach)Tj
2.6078 0 Td
(Company)Tj
4.1613 0 Td
(to)Tj
1.0261 0 Td
(do)Tj
1.2529 0 Td
(surveys)Tj
-34.3783 -1.3039 Td
(at)Tj
0.9524 0 Td
(Stillwater)Tj
3.9628 0 Td
(Cove.)Tj
2.4945 0 Td
(Hopkins)Tj
3.685 0 Td
(Marine)Tj
3.1578 0 Td
(Station)Tj
3.05 0 Td
(provided)Tj
3.8154 0 Td
(access)Tj
2.6476 0 Td
(to)Tj
1.0204 0 Td
(the)Tj
1.474 0 Td
(site)Tj
1.5704 0 Td
(within)Tj
2.812 0 Td
(their)Tj
2.1089 0 Td
(reserve.)Tj
3.2712 0 Td
(El)Tj
-36.0224 -1.2983 Td
(Sur)Tj
1.5817 0 Td
(Ranch)Tj
2.7666 0 Td
(allowed)Tj
3.2825 0 Td
(access)Tj
2.6362 0 Td
(to)Tj
1.0148 0 Td
(Andrew)Tj
3.4525 0 Td
(Molera.)Tj
3.3109 0 Td
(Permission)Tj
4.6941 0 Td
(to)Tj
1.0148 0 Td
(access)Tj
2.6362 0 Td
(sites)Tj
1.9219 0 Td
(located)Tj
3.0727 0 Td
(on)Tj
1.2643 0 Td
(Vandenberg)Tj
-32.6492 -1.2982 Td
(Air)Tj
1.542 0 Td
(Force)Tj
2.4718 0 Td
(Base)Tj
2.0182 0 Td
(was)Tj
1.7008 0 Td
(granted)Tj
3.2882 0 Td
(by)Tj
1.1792 0 Td
(the)Tj
1.474 0 Td
(U.S.)Tj
1.8822 0 Td
(Air)Tj
1.542 0 Td
(Force.)Tj
2.6986 0 Td
(Access)Tj
2.897 0 Td
(to)Tj
1.0204 0 Td
(Government)Tj
5.3348 0 Td
(Point)Tj
2.3981 0 Td
(was)Tj
1.7008 0 Td
(provided)Tj
-33.1481 -1.3039 Td
(by)Tj
1.1792 0 Td
(Bixby)Tj
2.4944 0 Td
(Management)Tj
5.5105 0 Td
(Inc.)Tj
1.7575 0 Td
(Access)Tj
2.897 0 Td
(to)Tj
1.0205 0 Td
(Alegria)Tj
3.1124 0 Td
(was)Tj
1.7008 0 Td
(approved)Tj
3.9741 0 Td
(annually)Tj
3.6567 0 Td
(by)Tj
1.1849 0 Td
(the)Tj
1.4683 0 Td
(Hollister)Tj
3.7077 0 Td
(Ranch)Tj
-33.664 -1.2983 Td
(Corporation.)Tj
5.4368 0 Td
(Sites)Tj
2.0466 0 Td
(not)Tj
1.5704 0 Td
(specifically)Tj
4.575 0 Td
(listed)Tj
2.3414 0 Td
(above)Tj
2.5455 0 Td
(as)Tj
1.0148 0 Td
(requiring)Tj
3.9685 0 Td
(permits)Tj
3.2825 0 Td
(were)Tj
2.1203 0 Td
(either)Tj
2.5342 0 Td
(on)Tj
1.2699 0 Td
(public)Tj
2.7099 0 Td
(land)Tj
-35.4158 -1.2983 Td
(or)Tj
1.0941 0 Td
(were)Tj
2.126 0 Td
(surveyed)Tj
3.7644 0 Td
(by)Tj
1.1792 0 Td
(the)Tj
1.474 0 Td
(organization)Tj
5.2781 0 Td
(in)Tj
1.0374 0 Td
(charge)Tj
2.863 0 Td
(of)Tj
1.0148 0 Td
(granting)Tj
3.617 0 Td
(access)Tj
2.6475 0 Td
(or)Tj
1.0999 0 Td
(permits)Tj
3.2768 0 Td
(\(e.g.,)Tj
2.126 0 Td
(National)Tj
-32.5982 -1.2982 Td
(Park)Tj
2.0806 0 Td
(Service\).)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 276.1511 cm
BT
/F11 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Study)Tj
2.6173 0 Td
(sites)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 259.1433 cm
BT
/F6 1 Tf
1 TL
-0.01 Tc
0 0 Td
(MARINe)Tj
3.8777 0 Td
(is)Tj
0.8277 0 Td
(a)Tj
0.6463 0 Td
(consortium)Tj
4.7849 0 Td
(of)Tj
0.9978 0 Td
(18)Tj
1.1508 0 Td
(groups)Tj
2.931 0 Td
(\(including)Tj
4.3143 0 Td
(state,)Tj
2.2224 0 Td
(federal,)Tj
3.1067 0 Td
(university,)Tj
4.3484 0 Td
(and)Tj
1.7064 0 Td
(private)Tj
2.9367 0 Td
(organiza-)Tj
-33.8511 -1.2982 Td
(tions\))Tj
2.5058 0 Td
(that)Tj
1.7518 0 Td
(conduct)Tj
3.4355 0 Td
(coordinated)Tj
4.989 0 Td
(annual)Tj
2.9253 0 Td
(monitoring)Tj
4.7565 0 Td
(of)Tj
1.0035 0 Td
(intertidal)Tj
3.8437 0 Td
(community)Tj
4.8359 0 Td
(parameters)Tj
4.6091 0 Td
(at)Tj
0.9411 0 Td
(over)Tj
-35.5972 -1.3039 Td
(130)Tj
1.6214 0 Td
(sites)Tj
1.8935 0 Td
(in)Tj
1.0148 0 Td
(four)Tj
1.8822 0 Td
(US)Tj
1.4173 0 Td
(states.)Tj
2.5738 0 Td
(By)Tj
1.2303 0 Td
(combining)Tj
4.5127 0 Td
(data)Tj
1.8879 0 Td
(from)Tj
2.1713 0 Td
(MARINe)Tj
3.8777 0 Td
(groups)Tj
2.9254 0 Td
(with)Tj
1.9672 0 Td
(data)Tj
1.8822 0 Td
(from)Tj
2.177 0 Td
(the)Tj
1.4457 0 Td
(Univer-)Tj
-34.4804 -1.2983 Td
(sity)Tj
1.576 0 Td
(of)Tj
1.0035 0 Td
(British)Tj
2.8516 0 Td
(Columbia,)Tj
4.3597 0 Td
(we)Tj
1.3152 0 Td
(present)Tj
3.1238 0 Td
(data)Tj
1.8878 0 Td
(on)Tj
1.2643 0 Td
(populations)Tj
4.8869 0 Td
(of)Tj
/F10 1 Tf
1.0034 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9808 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
3.9798 0 Td
(at)Tj
0.9411 0 Td
(90)Tj
1.1566 0 Td
(rocky)Tj
2.4434 0 Td
(intertidal)Tj
-32.7739 -1.2983 Td
(sites)Tj
1.8992 0 Td
(spanning)Tj
3.8607 0 Td
(the)Tj
1.4513 0 Td
(North)Tj
2.6306 0 Td
(American)Tj
4.1442 0 Td
(Pacific)Tj
2.8403 0 Td
(coast,)Tj
2.4378 0 Td
(from)Tj
2.177 0 Td
(southeast)Tj
3.9061 0 Td
(Alaska)Tj
2.8403 0 Td
(to)Tj
1.0148 0 Td
(San)Tj
1.6611 0 Td
(Diego)Tj
2.5852 0 Td
(County,)Tj
-33.4486 -1.3039 Td
(California)Tj
4.1839 0 Td
(\()Tj
0.83 0.64 0.02 0 k
(Fig)Tj
1.7971 0 Td
(1)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9071 0 Td
(S1AS1H)Tj
4.0535 0 Td
(Fig)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.6781 0 Td
(S1)Tj
1.1622 0 Td
(Table)Tj
0 g
(\).)Tj
2.965 0 Td
(Physical)Tj
3.4469 0 Td
(site)Tj
1.5421 0 Td
(characteristics)Tj
5.8166 0 Td
(such)Tj
2.0353 0 Td
(as)Tj
1.0035 0 Td
(rock)Tj
1.9955 0 Td
(type,)Tj
2.109 0 Td
(bench)Tj
-34.6958 -1.2982 Td
(slope)Tj
2.245 0 Td
(and)Tj
1.7121 0 Td
(size,)Tj
1.8765 0 Td
(wave)Tj
2.194 0 Td
(exposure,)Tj
3.9912 0 Td
(temperature)Tj
5.0626 0 Td
(\(both)Tj
2.3698 0 Td
(air)Tj
1.2699 0 Td
(and)Tj
1.7178 0 Td
(water\),)Tj
2.9536 0 Td
(and)Tj
1.7122 0 Td
(sand)Tj
2.0636 0 Td
(influence)Tj
3.8664 0 Td
(vary)Tj
1.9105 0 Td
(sub-)Tj
-34.9452 -1.2983 Td
(stantially)Tj
3.7473 0 Td
(across)Tj
2.6419 0 Td
(this)Tj
1.6498 0 Td
(broad)Tj
2.5398 0 Td
(stretch)Tj
2.88 0 Td
(of)Tj
1.0034 0 Td
(coastline,)Tj
3.8948 0 Td
(but)Tj
1.5364 0 Td
(all)Tj
1.1281 0 Td
(sites)Tj
1.8993 0 Td
(contained)Tj
4.1442 0 Td
(appropriate)Tj
4.8132 0 Td
(habitat)Tj
2.9423 0 Td
(for)Tj
/F10 1 Tf
1.3663 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
-36.1868 -1.3039 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
(.)Tj
4.1952 0 Td
(Detailed)Tj
3.5319 0 Td
(site)Tj
1.5421 0 Td
(descriptions,)Tj
5.244 0 Td
(including)Tj
3.9855 0 Td
(monitoring)Tj
4.7622 0 Td
(duration)Tj
3.651 0 Td
(and)Tj
1.7121 0 Td
(frequency,)Tj
4.354 0 Td
(photos,)Tj
-32.978 -1.2983 Td
(and)Tj
1.7121 0 Td
(trends)Tj
2.7155 0 Td
(for)Tj
1.3663 0 Td
(other)Tj
2.3188 0 Td
(species)Tj
2.9536 0 Td
(monitored,)Tj
4.6545 0 Td
(can)Tj
1.6101 0 Td
(be)Tj
1.1395 0 Td
(found)Tj
2.5852 0 Td
(at)Tj
0.83 0.64 0.02 0 k
0.9411 0 Td
(pacificrockyintertidal.or)Tj
9.513 0 Td
(g)Tj
0 g
(.)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 112.9889 cm
BT
/F11 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Abundance)Tj
4.989 0 Td
(and)Tj
1.7905 0 Td
(size)Tj
1.767 0 Td
(structure:)Tj
4.2898 0 Td
(Long-term)Tj
4.6819 0 Td
(monitoring)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 95.9811 cm
BT
/F10 1 Tf
1 TL
-0.004 Tc
0 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
/F10 1 Tf
0.9921 0 Td
[()4()13()18(\n)20()6()7()2()-4(\r)]TJ
/F6 1 Tf
4.0308 0 Td
(were)Tj
2.126 0 Td
(counted,)Tj
3.702 0 Td
(measured,)Tj
4.3313 0 Td
(and)Tj
1.7291 0 Td
(\(beginning)Tj
4.6035 0 Td
(in)Tj
1.0318 0 Td
(2013\))Tj
2.4661 0 Td
(assigned)Tj
3.6057 0 Td
(to)Tj
1.0261 0 Td
(disease)Tj
3.016 0 Td
(categories)Tj
-32.6605 -1.2982 Td
(annually)Tj
3.6566 0 Td
(or)Tj
1.0999 0 Td
(semiannually)Tj
5.5331 0 Td
(at)Tj
0.9525 0 Td
(90)Tj
1.1679 0 Td
(long-term)Tj
4.2519 0 Td
(monitoring)Tj
4.8189 0 Td
(\(LTM\))Tj
2.9423 0 Td
(sites)Tj
1.9332 0 Td
(in)Tj
1.0318 0 Td
(Alaska,)Tj
3.1011 0 Td
(Washington,)Tj
ET
Q
q
1 j
1 J
0 w
576 737.1 m
36 737.1 l
36 737.6 l
576 737.6 l
f*
36 741.2598 107.1496 23.6976 re
W* n
q
107.0929 0 0 23.6409 36 741.3165 cm
q
/I0 Do
Q
Q
Q
q
0 0 612 792 re
W* n
1 j
1 J
0 w
7.9999 0 0 7.9999 397.0771 745.7952 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(Large-scale)Tj
5.3504 0 Td
(impacts)Tj
3.6071 0 Td
(of)Tj
1.0417 0 Td
(SSWD)Tj
3.1961 0 Td
(on)Tj
1.3111 0 Td
(interti)Tj
2.3882 0 Td
(dal)Tj
1.5236 0 Td
(sea)Tj
1.8071 0 Td
(stars)Tj
ET
Q
q
1 j
1 J
0 w
36 48.0002 m
576 48.0002 l
576 47.5002 l
36 47.5002 l
f*
0.83 0.64 0.02 0 k
81.5811 34.9228 m
237.8835 34.9228 l
h
f*
0 g
7.9999 0 0 7.9999 36 36 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(PLOS)Tj
2.8559 0 Td
(ONE)Tj
2.3669 0 Td
(|)Tj
0.83 0.64 0.02 0 k
0.4748 0 Td
(https://doi.or)Tj
5.4142 0 Td
(g/10.137)Tj
3.8198 0 Td
(1/journal.po)Tj
5.1236 0 Td
(ne.01928)Tj
4.089 0 Td
(70)Tj
0 g
2.0906 0 Td
(March)Tj
2.9622 0 Td
(20,)Tj
1.5874 0 Td
(2018)Tj
34.3562 0 Td
(4)Tj
0.7654 0 Td
(/)Tj
ET
endstream
endobj
122 0 obj
<>stream
endstream
endobj
123 0 obj
<>stream
endstream
endobj
124 0 obj
<>stream
endstream
endobj
125 0 obj
<>stream
endstream
endobj
126 0 obj
<>stream
endstream
endobj
127 0 obj
<>stream
endstream
endobj
128 0 obj
<>stream
BT
66.402 0 Td
(21)Tj
ET
Q
endstream
endobj
129 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 12 0 R/Annots 131 0 R/Contents 132 0 R/TrimBox[0 0 612 792]>>
endobj
130 0 obj
<>stream
xImU{!r!%Df-DbYH(P*HHapFb[Ilcc^9^v}9cm-,z_oc8{ENG#p?G#p?G#po??^1eGP7F1flCWAxf~G}TpC]o@nO4kJ1S"P7zYI=JZ1k썵@@9LZ'Schu_}Zn>MDe[Еex鲁l?0K[N9S[Ʈ`KGݑپp3
oKh
mNeZ8msi>vNvvMkaRf2?Zbi\ !Ï춸m>J]t`-W1`AKƻtL#
5p; LlxєߎO< 9bomw5i a.p]v6?·Τ]b(|2)5eö:(4@'!-D.fZ)Y